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Gonocephalus liogaster (GÜNTHER, 1872)

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Higher TaxaAgamidae (Draconinae), Sauria, Iguania, Squamata (lizards)
Subspecies 
Common NamesE: Tropical Forest Dragon, Blue-eyed Anglehead Lizard, Orange-ringed Anglehead Lizard
G: Blauaugen-Winkelkopfagame 
SynonymTiaris liogaster GÜNTHER 1872: 592
Gonyocephalus liogaster — BOULENGER 1885: 286
Goniocephalus herveyi BOULENGER 1887
Gonyocephalus liogaster — DE ROOIJ 1915: 103
Gonyocephalus herveyi — DE ROOIJ 1915: 104
Gonocephalus liogaster — WERMUTH 1967: 60
Gonocephalus liogaster — MANTHEY & GROSSMANN 1997: 187
Gonocephalus liogaster — MANTHEY & SCHUSTER 1999: 64
Gonocephalus liogaster — GRISMER 2011
Gonocephalus liogaster — GRISMER & QUAH 2019 
DistributionIndonesia (Sumatra, Borneo, Natuna Island), West Malaysia

Type locality: Sarawak, Borneo  
Reproductionoviparous 
TypesSyntypes: BMNH (NHMUK) 1946.8.14.79, male, coll. Everett; NHMUK 1946.8.27.16, male, Borneo, (coll. Everett, donation Cutter?); NHMUK 1946.8.27.17, male, coll. Bleeker, donation; NHMUK 1946.8.13.86, juvenile, Borneo, coll. A.R. Wallace; NHMUK 1946.8.14.82, coll. G. Doria. 
DiagnosisDescription: Snout longer than the diameter of the orbit; canthus rostralis
sharp, projecting; supraciliary border normally raised; tympanum smaller than the eye-opening. Head-scales small, keeled, largest on the supraorbital region; some enlarged tubercles above and behind the ear; nine to eleven upper and nine to twelve lower labials. Gular sac moderate, without serrated edge and covered with small smooth scales. Body strongly compressed; nuchal and dorsal crests continuous, very high, beginning at the occiput, and formed of long narrow scales, longer than the snout, covered at the base with three rows of smaller keeled ones, the superior row twice as high as the lower. In females the crest is much lower. Dorsal scales very small, smooth or feebly keeled, the points directed upwards, with a few scattered enlarged ones; ventral scales larger than dorsals, smooth. Tail compressed, with a serrated edge and covered with smooth or feebly keeled scales, the inferior largest and strongly keeled; its length more than twice that of head and body. Limbs long, the hind limb reaches the tip of the snout or beyond, covered with unequal slightly keeled scales; fourth finger a little longer than third (de Rooij 1915: 103).

Coloration: Olive or brown above, uniform or with dark transverse bands. Limbs banded with dark brown; tail with black and light annuli. Lower parts brownish; gular fold black; gular sac black-spotted, the spots arranged in oblique series, or entirely black (de Rooij 1915: 103).

Size: Length of head and body 141 mm.; tail 315 mm (de Rooij 1915: 103). 
CommentSynonymy mostly after WERMUTH 1967. Has been reported from Vietnam (Malacca) but this has not been confirmed (V. Bobrov, pers. comm.). 
References
  • ARIFIN, UMILAELA; DJOKO T. ISKANDAR and ROSITA ELIANUR 2011. Herpetofauna Diversity Of Karimata Island-Indonesia. Proceedings of the Conference “Biology of the Amphibians in the Sunda Region, South-east Asia”. Organized by Universiti Malaysia Sarawak, 28–30 September 2009. pp:1–7. Edited by Indraneil Das et al.. Institute of Biodiversity and Environmental Conse - get paper here
  • Auliya, M. 2006. Taxonomy, Life History, and conservation of giant reptiles in west Kalimantan. Natur und Tier Verlag, Münster, 432 pp. - get paper here
  • Boulenger, G.A. 1885. Catalogue of the lizards in the British Museum (Nat. Hist.) I. Geckonidae, Eublepharidae, Uroplatidae, Pygopodidae, Agamidae. London: 450 pp. - get paper here
  • Campbell, P. et al. 2011. FHF snapshot. HerpNation 2.4 (7): 18-22 - get paper here
  • Das , I. 2004. Lizards of Borneo. Natural History Publications, Kota Kinabalu, Borneo
  • Das, Indraneil; Jye Wen Wong 2021. Predation on Gonocephalus liogaster (Agamidae) by Ptyas carinata (Colubridae) in Sarawak, Borneo. Herpetology Notes 14: 349-351 - get paper here
  • de Rooij, N. de 1915. The Reptiles of the Indo-Australian Archipelago. I. Lacertilia, Chelonia, Emydosauria. Leiden (E. J. Brill), xiv + 384 pp. - get paper here
  • Dring J C M 1979. Amphibians and reptiles from northern Trengganu, Malaysia, with descriptions of two new geckos: Cnemaspis and Cyrtodactylus. Bulletin of the British Museum (NATURAL HISTORY) ZOOLOGY 34 (5): 181-241 - get paper here
  • GRISMER, L. LEE; & EVAN S. H. QUAH 2019. An updated and annotated checklist of the lizards of Peninsular Malaysia, Singapore, and their adjacent archipelagos. Zootaxa 4545 (2): 230–248 - get paper here
  • Grismer, L.L. 2011. Lizards of Peninsular Malaysia, Singapore and their adjacent archipelagos. Edition Chimaira, Frankfurt, 728 pp. [review in Herp. Rev. 43: 155] - get paper here
  • Günther, A. 1872. On the reptiles and amphibians of Borneo. Proc. Zool. Soc. London 1872: 586-600. - get paper here
  • Herlambang, A. E., Riyanto, A., Munir, M., Hamidy, A., Kimura, K., Eto, K., & Mumpuni, M. 2022. AFTER 16 YEARS: AN UPDATE CHECKLIST OF HERPETOFAUNA ON THE NATUNA ISLANDS, INDONESIA. TREUBIA, 49(2), 67-84 - get paper here
  • Janzen, Peter 2022. Herpetofauna in Sarawak, East Malaysia (Borneo) – Part 2 Reptiles. Sauria 44 (3): 3-28
  • Leo S, Suherman M, Permatasari A, Suganda D, Zulamri, Winarni NL. 2020. Herpetofauna diversity in Zamrud National Park, Indonesia: baseline checklist for a Sumatra peat swamp forest ecosystem. Amphibian & Reptile Conservation 14(2) [General Section]: 250–263 (e249) - get paper here
  • Manthey U 2010. Agamid Lizards of Southern Asia. Draconinae 2 -Leiolepidinae. Edition Chimaira, Terralog 7b, Frankfurt, 168 pp.
  • Manthey, U. & Grossmann, W. 1997. Amphibien & Reptilien Südostasiens. Natur und Tier Verlag (Münster), 512 pp. - get paper here
  • Manthey,U. & SCHUSTER,N. 1999. Agamen, 2. Aufl. Natur und Tier Verlag (Münster), 120 pp. - get paper here
  • NUR-AMALINA M.I.; AZHARI, M., NORSHAQINAH, A., NOR AZRIN, N.A., SHUKOR, M.N., AISAH, M.S., AMIRRUDIN, A., GRISMER, L.L. and NORHAYATI, A. 2017. SPECIES COMPOSITION OF AMPHIBIANS AND REPTILES IN TEMBAT FOREST RESERVE, HULU TERENGGANU, TERENGGANU, PENINSULAR MALAYSIA. Malays. Appl. Biol. 46 (4): 119–129 - get paper here
  • Sumarli, Alex X.; L. Lee Grismer, Shahrul Anuar, Mohd Abdul Muin and Evan S.H. Quah 2015. First report on the amphibians and reptiles of a remote mountain, Gunung Tebu in northeastern Peninsular Malaysia. Check List 11 (4): 1679 - get paper here
  • TEYNIÉ, ALEXANDRE; PATRICK DAVID, & ANNEMARIE OHLER 2010. Note on a collection of Amphibians and Reptiles from Western Sumatra (Indonesia), with the description of a new species of the genus Bufo. Zootaxa 2416: 1–43 - get paper here
  • Wong, A., Makbul, N. S., & Yambun, P. 2021. Diversity of Herpetofauna in Kadamaian, Kota Belud, Sabah, Malaysia. Journal of Tropical Biology & Conservation (JTBC), 18, 31-42 - get paper here
 
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