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Gonocephalus sophiae (GRAY, 1845)

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Higher TaxaAgamidae (Draconinae), Sauria, Iguania, Squamata (lizards)
Common NamesE: Negros Forest Dragon
G: Philippinische Winkelkopfagame 
SynonymTiaris sophiae GRAY 1845: 240
Tiaris bellii — GRAY 1845: 239 (non Lophyrus bellii DUMERIL & BIBRON)
Tiaris petersii GÜNTHER 1868: 136
Gonyocephalus sophiae — BOULENGER 1885: 288
Gonyocephalus sophiae — TAYLOR 1922: 133
Gonocephalus sophiae — WERMUTH 1967: 62
Gonocephalus sophiae – ALCALA 1986: 86
Gonocephalus sophiae — GAULKE 2001: 7
Gonocephalus cf. sophiaei — BINADAY et al. 2017 
DistributionPhilippines (Negros, Mindanao, Panay, Masbate, Luzon, Cebu)

Type locality: Philippines.  
TypesHolotype: BMNH 1946.8.27.11-12, 
CommentSynonymy partly after WERMUTH 1967. 
  • Alcala, A.C. 1986. Guide to Philippine flora and fauna. Vol. X. Amphibians and reptiles. Manila, Natural Resources Management Center, Ministry of Natural Resources, University of the Philippines: i-xiv + 1-195
  • Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here
  • Binaday, Jake Wilson B.; Ace Kevin S. Amarga, Ernesto S. Barrameda Jr., and Bobet Jan M. Bonagua 2017. Amphibians and Reptiles in the Vicinity of Bulusan Lake, Bulusan Volcano Natural Park, Sorsogon, Philippines. Philippine Journal of Science 146 (3): 339-351
  • Boulenger, G.A. 1885. Catalogue of the lizards in the British Museum (Nat. Hist.) I. Geckonidae, Eublepharidae, Uroplatidae, Pygopodidae, Agamidae. London: 450 pp. - get paper here
  • Gaulke, M. 2001. Die Herpetofauna von Sibaliw (Panay), einem der letzten Tieflandregenwaldgebiete der West-Visayas, Philippinen. Teil III. Echsen und Diskussion. Herpetofauna 23 (132): 5-18 - get paper here
  • Gaulke, M. 2011. The herpetofauna of Panay Island, Philippines. Edition Chimaira, 390 pp.
  • Gaulke, M. & A. D. Demegillo 2008. Der Mabitang (Varanus mabitang), ein Früchte fressender Großwaran von den Philippinen. Reptilia (Münster) 13 (71): 67-74 - get paper here
  • Gaulke, M. & Demegillo, A. 2005. Biology and variability of the Negros Forest Dragon Gonocephalus sophiae. Reptilia (GB) (42): 16-23 - get paper here
  • Gaulke, M. & Demegillo, A. 2005. Zur Variabilität und Biologie der philippinischen Winkelkopfagame, Gonocephalus sophiae. Reptilia (Münster) 10 (51): 32-40 - get paper here
  • Gojo-Cruz, Paul Henric P. and Leticia E. Afuang 2018. The Zoogeographic Significance of Caraballo Mountain Range, Luzon Island, Philippines With Focus on the Biogeography of Luzon’s Herpetofauna. Philippine Journal of Science 147 (3): 393-409 - get paper here
  • Gray, J. E. 1845. Catalogue of the specimens of lizards in the collection of the British Museum. Trustees of die British Museum/Edward Newman, London: xxvii + 289 pp. - get paper here
  • Günther,A. 1868. Zool. Rec. (Amph. Rept.) 1867: 136
  • Lagat, Ronaldo D. and Rubie M. Causaren 2019. Initial terrestrial vertebrate diversity assessment in upland Cavite, Philippines. Philippine Journal of Systematic Biology 12 (2): 70-91 - get paper here
  • Manthey U 2010. Agamid Lizards of Southern Asia. Draconinae 2 -Leiolepidinae. Edition Chimaira, Terralog 7b, Frankfurt, 168 pp.
  • MCLEOD, David S.; Cameron D. SILER, Arvin C. DIESMOS, Mae L. DIESMOS, Vhon S. GARCIA, Angela O. ARKONCEO, Kelvin L. BALAQUIT, Charlene C. UY, Mariden M. VILLASERAN, Earle C. YARRA, Rafe M. BROWN 2011. Amphibians and Reptiles of Luzon Island, V: The Herpetofauna of Angat Dam Watershed, Bulacan Province, Luzon Island, Philippines. Asian Herpetological Research 2 (4): 177–198 - get paper here
  • Siler, C.D.; L.J. Welton; J.M. Siler; J. Brown; A. Bucol; A.C. Diesmos; R. Brown. 2011. Amphibians and Reptiles, Luzon Island, Aurora Province and Aurora Memorial National Park, Northern Philippines: New island distribution records. Check List 7 (2):182-195 - get paper here
  • Siler, Cameron D. and Rafe M. Brown 2010. Phylogeny-based Species Delimitation in Philippine Slender Skinks (Reptilia: Squamata: Scincidae: Brachymeles): Taxonomic Revision of Pentadactyl Species Groups and Description of Three New Species. Herpetological Monographs 24 (1): 1-54 - get paper here
  • SUPSUP, Christian E.; Nevong M. PUNA, Augusto A. ASIS, Bernard R. REDOBLADO, Maria Fatima G. PANAGUINIT, Faith M. GUINTO, Edmund B. RICO, Arvin C. DIESMOS, Rafe M. BROWN and Neil Aldrin D. MALLARI 2016. Amphibians and Reptiles of Cebu, Philippines: The Poorly Understood Herpetofauna of an Island with Very Little Remaining Natural Habitat. Asian Herpetological Research 2016, 7(3): 151–179 DOI: 10.16373/j.cnki.ahr.150049 - get paper here
  • Taylor, E.H. 1922. The lizards of the Philippine Islands. Department of Agriculture and Natural Resources, Bureau of Science, Government of the Philippine Islands, Manila, Publication no. 17: 269 pp. - get paper here
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