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Harpesaurus borneensis (MERTENS, 1924)

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Higher TaxaAgamidae (Draconinae), Sauria, Iguania, Squamata (lizards)
Subspecies 
Common Names 
SynonymHylagama borneensis MERTENS 1924
Harpesaurus thescelorhinos KING 1978
Hylagama borneensis — WERMUTH 1967: 65
Harpesaurus borneensis — MANTHEY & GROSSMANN 1997: 192
Hylagama borneensis — MANTHEY & SCHUSTER 1999: 70
Harpesaurus borneensis — BARTS & WILMS 2003
Harpesaurus borneensis — MANTHEY 2010 
DistributionIndonesia (Borneo, Sarawak, Kalimantan)

thescelorhinos: MALAYSIA (Sarawak, Borneo)

Type locality: Central Borneo, probably Kapoeas area.  
Reproductionovoviviparous 
TypesHolotype: SMNS 4596, adult female. Moody (1984) considered the holotype of Hylagama borneensis as lost and (erroneously) designated FMNH 138196 erroneously as a lectotype (must be neotype) to replace the lost holotype (designation invalid as holotype still extant).
Holotype: adult male, FMNH 131540; adult female, paratype, FMNH 138196 [thescelorhinos] 
Diagnosis 
CommentMANTHEY & SCHUSTER 1999 note that there are no external differences to Harpesaurus but they still list this species as Hylagama.

Abundance: Rare. This is one of the species called 'lost' and 'rediscovered' by Lindken et al. 2024. 
EtymologyNamed after its distribution on Borneo (Indonesia). 
References
  • Barts, M. & Wilms, T. 2003. Die Agamen der Welt. Draco 4 (14): 4-23 - get paper here
  • Das , I. 2004. Lizards of Borneo. Natural History Publications, Kota Kinabalu, Borneo
  • Das, Indraneil; Wong Jye Wen & Veronica Martin 2019. Harpesaurus borneensis (Mertens, 1924). Sauria 41 (2): 1-3 - get paper here
  • King F W 1978. A new Bornean lizard of the genus Harpesaurus. Sarawak Museum Journal 26(No. 47) 1978: 205-209.
  • Lardner, Björn; Sing Yun Chin, and Ulrich Manthey 2010. NOTES ON Harpesaurus borneensis (MERTENS 1924), A LIVE BEARING AGAMID LIZARD FROM THE LOWLANDS OF BORNEO. Russ. J. Herpetol. 17 (3): 231 – 235 - get paper here
  • Lindken T.; Anderson, C. V., Ariano-Sánchez, D., Barki, G., Biggs, C., Bowles, P., Chaitanya, R., Cronin, D. T., Jähnig, S. C., Jeschke, J. M., Kennerley, R. J., Lacher, T. E. Jr., Luedtke, J. A., Liu, C., Long, B., Mallon, D., Martin, G. M., Meiri, 2024. What factors influence the rediscovery of lost tetrapod species? Global Change Biology, 30: 1-18 - get paper here
  • Manthey U 2010. Agamid Lizards of Southern Asia. Draconinae 2 -Leiolepidinae. Edition Chimaira, Terralog 7b, Frankfurt, 168 pp.
  • Manthey, U. & Grossmann, W. 1997. Amphibien & Reptilien Südostasiens. Natur und Tier Verlag (Münster), 512 pp. - get paper here
  • Manthey,U. & SCHUSTER,N. 1999. Agamen, 2. Aufl. Natur und Tier Verlag (Münster), 120 pp. - get paper here
  • Mertens, R. 1924. Über einige Reptilien aus Borneo. Zool. Anz. 60: 155-159
  • Schlüter, A. & Hallermann, J. 1997. The Type Specimens in the Herpetological Collection of the Staatliches Museum für Naturkunde in Stuttgart. Stuttgarter Beitr. Naturk. Ser. A (553): 1-15 - get paper here
 
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