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Harpesaurus tricinctus (DUMÉRIL, 1851)

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Higher TaxaAgamidae (Draconinae), Sauria, Iguania, Squamata (lizards)
Subspecies 
Common NamesE: Java Nose-horned Lizard 
SynonymArpephorus tricinctus DUMÉRIL in DUMÉRIL & DUMÉRIL 1851: 93
Harpesaurus tricinctus — BOULENGER 1885: 279
Harpesaurus tricinctus — DE ROOIJ 1915: 96
Harpesaurus tricinctus — MANTHEY & GROSSMANN 1997: 193
Harpesaurus tricinctus — MANTHEY & SCHUSTER 1999: 66
Harpesaurus tricinctus — MANTHEY 2010
Harpesaurus tricinctus — PUTRA et al. 2020 
DistributionIndonesia (Java, Borneo ?),
Malaysia (Borneo ?, Sarawak)

Type locality: Java  
Reproductionoviparous 
TypesHolotype: MNHN-RA 0623 
DiagnosisDescription: Rostral appendage longer than the head, formed of a compressed falciform scale curved upwards, surrounded at the base by a few large scales. Head with small scales, slightly tubercular; a large prominent, triangular scale on the snout; canthus rostralis a serrated ridge. Gular scales tubercular. Body compressed; dorsal crest a low serrated ridge; dorsal scales smooth, equal, in regular transverse series; ventral scales keeled. Tail compressed, with a crest above, slightly higher than the dorsal, covered with keeled scales, below with a double series of spinose scales. Limbs covered with keeled scales (de Rooij 1915: 96).

Coloration: Brown above, with three broad transverse yellow bands on the body, the anterior the narrowest, on the shoulders, the others descending on the flanks to the belly, where they do not meet (de Rooij 1915: 96).

Size: Length of head and body (without rostral appendage) 64mm.; tail 83 mm.; rostral appendage 21 mm (de Rooij 1915: 96). 
CommentType Species: Arpephorus tricinctus DUMÉRIL in DUMÉRIL & DUMÉRIL 1851 is the type species of the genus Harpesaurus BOULENGER 1885. Arpephorus is preoccupied by Arpephorus STEPHENS 1829 (Coleoptera).

The most striking feature of the genus is the presence, either in males only or in both sexes, of one or two prominent rostral appendages. 
References
  • Boulenger, G.A. 1885. Catalogue of the lizards in the British Museum (Nat. Hist.) I. Geckonidae, Eublepharidae, Uroplatidae, Pygopodidae, Agamidae. London: 450 pp. - get paper here
  • de Rooij, N. de 1915. The Reptiles of the Indo-Australian Archipelago. I. Lacertilia, Chelonia, Emydosauria. Leiden (E. J. Brill), xiv + 384 pp. - get paper here
  • Duméril, A.M.C. & A. H. A. Duméril 1851. Catalogue méthodique de la collection des reptiles du Muséum d'Histoire Naturelle de Paris. Gide et Baudry/Roret, Paris, 224 pp.
  • Manthey U 2010. Agamid Lizards of Southern Asia. Draconinae 2 -Leiolepidinae. Edition Chimaira, Terralog 7b, Frankfurt, 168 pp.
  • Manthey, U. & Grossmann, W. 1997. Amphibien & Reptilien Südostasiens. Natur und Tier Verlag (Münster), 512 pp. - get paper here
  • Manthey,U. & SCHUSTER,N. 1999. Agamen, 2. Aufl. Natur und Tier Verlag (Münster), 120 pp. - get paper here
  • Putra, C.A. et al. 2020. Rediscovery of Modigliani’s nose-horned lizard, Harpesaurus modiglianii Vinciguerra, 1933 (Reptilia: Agamidae) after 129 years without any observation. TAPROBANICA 9 (1): 3-11 - get paper here
 
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