Hebius maximus (MALNATE, 1962)

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Higher Taxa	Colubridae (Natricinae), Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies
Common Names	E: Western China Keelback Chinese: 华西腹链蛇 (Huá Xī Fù Liàn Shé)
Synonym	Amphiesma sauteri maximus MALNATE 1962: 272 Natrix sauteri — POPE 1935: 125 (in part) Natrix sauteri — WU 1985: 209 (in part) Amphiesma sauteri — INGER et al. 1990: 20 Amphiesma sauteri — ZHAO & ADLER 1993: 228 (in part) Amphiesma sauteri — ZHAO et al. 1998: 80 (in part) Amphiesma sauteri — ZHAO 2003: 140 Amphiesma sauteri — DAVID et al. 2007: 60 (in part) Amphiesma sauteri — WALLACH et al. 2014: 33 (in part). Hebius sauteri — GUO et al. 2014: 428 (in part) Hebius sauteri — DAVID et al. 2021 (in part) Hebius maximus — LI et al. 2022
Distribution	SW China (Chongqing: Youyang Tujia and Miao Autonomous County, Xiushan Tujia and Miao Autonomous County, Sichuan: Mabian County, Pingshan County, Yingjing County, Hongya County, Dujiangyan City, Mianyang City, Leshan City, Guizhou: Qixingguan District, Bijie City) Type locality: Hsaioyangchi, Szechwan, China (= Xiaoyangxi, Mabian Yi Autonomous County, Leshan City, Sichuan Province, China)
Reproduction	oviparous
Types	Holotype: FMNH 18796 (formerly CHNM 18796), a 457 mm adult female specimen (Fig. 7). Paratypes. FMNH 18797, USNM 71570, USNM 84362.
Diagnosis	Unfortunately we had to temporarily remove additional information as this was scraped by multiple AI companies who sell that data to their customers. These details, e.g. detailed descriptions or comparisons (about 6441 characters), are available for collaborators and contributors. Please contact us for details.
Comment	Synonymy: after Li et al. 2022, who suggested to elevate maximus to full species level, first in a preprint, then in a peer-reviewed paper.
Etymology	Hebius maximus comb. nov. is named for its highest count of ventral scales among three previous subspecies of H. sauteri (Malnate 1962).
References	David, P., Bain, R. H., Nguyen Quang Truong, Orlov, N. L., Vogel, G., Vu Ngoc Thanh & T. Ziegler 2007. A new species of the natricine snake genus Amphiesma from the Indochinese Region (Squamata: Colubridae: Natricinae). Zootaxa 1462: 41-60 - get paper here DAVID, PATRICK; GERNOT VOGEL, TRUONG QUANG NGUYEN, NIKOLAI L. ORLOV, OLIVIER S. G. PAUWELS, ALEXANDRE TEYNIÉ, THOMAS ZIEGLER 2021. A revision of the dark-bellied, stream-dwelling snakes of the genus Hebius (Reptilia: Squamata: Natricidae) with the description of a new species from China, Vietnam and Thailand. Zootaxa 4911: 1–61 - get paper here Inger R F; Zhao E; Bradley Shaffer H; Guanfu Wu 1990. Report on a collection of amphibians and reptiles from Sichuan, China. Fieldiana: Zoology (No. 58) : i-iii, 1-24 - get paper here Li M-L, Ren J-L, Huang J-J, Lyu Z-T, Qi S, Jiang K, Wang Y-Y, Li J-T 2022. On the validity of Hebius sauteri maximus (Malnate, 1962) (Squamata, Natricidae), with the redescription of H. maximus comb. nov. and H. sauteri (Boulenger, 1909). Herpetozoa 35: 265-282 - get paper here Li, M.L., Ren, J.L., Huang, J.J., Lyu, Z.T., Qi, S., Jiang, K., Wang, Y.Y. and Li, J.T., 2022. On the validity of Hebius sauteri maximus (Squamata, Natricidae), with redescription of H. maximus comb. nov. and H. sauteri from China. ARPHA Preprints, 3, p.e94937 - get paper here Malnate, E.V. 1962. The relationships of five species of the Asiatic natricine snake genus Amphiesma. Proc. Acad. Nat. Sci. Philadelphia, 114, 251–299. - get paper here Malnate, Edmond V. 1960. Systematic division and evolution of the colubrid snake genus Natrix, with comments on the subfamily Natricinae. Proc. Acad. Nat. Sci. Philadelphia, 112 (3): 41-71. - get paper here Pope, CLIFFORD H. 1935. The Reptiles of China. Turtes, Crocodilians, Snakes, Lizards. Amer. Mus. Nat. Hist., New York, Nat. Hist. Central Asia, 10: lii, 1-604 - get paper here Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp. Wu Lu, Li Te-chun, & Liu Chi-ch'en 1985. Kuei-chou p'a hsing lei chih [Reptiles of Kweichow Province]. [People's Press], Kuei-yang 436 pp. Zhao E. M. 1998. Fauna Sinica, Reptilia, Vol. 3, Squamata: Serpentes. Science Press (In Chinese) Zhao, E. (ed.) 2003. Coloured atlas of Sichuan reptiles [in Chinese]. Beijing, China Forestry Publishing House Zhao,E. & Adler,K. 1993. Herpetology of China. SSAR, Oxford/Ohio, 1-522
External links	National Center for Biotechnology Information https://www.facebook.com/groups/stshc/posts/527032582854678/ Google images

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