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Hebius popei (SCHMIDT, 1925)

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Higher TaxaColubridae (Natricinae), Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies 
Common NamesE: Pope's Keelback
Chinese: 坡普腹链蛇 
SynonymNatrix popei SCHMIDT 1925: 3
Natrix vibakari popei — MELL 1931 [1929]
Natrix popei — GLASS 1946
Amphiesma popei MALNATE 1960
Amphiesma popei — DAVID et al. 2007
Hebius popei — GUO et al. 2014
Amphiesma popei — WALLACH et al. 2014: 32
Hebius popei — DAVID et al. 2021 
DistributionChina (Hainan, west to Hunan and Yunnan), Vietnam

Type locality: "Nodoa, Hainan, China."  
Reproductionoviparous 
TypesHolotype: AMNH 27763, adult male (C.H. Pope, Dec. 1922-July 1923). 
DiagnosisDIAGNOSIS. Closely allied to Natrix vibakari and to Natrix sauteri of Formosa; maxillary teeth 20, the last gradually enlarged; anal divided; scales in nineteen rows; one or two anterior temporals; eight upper labials; fourth and fifth entering the eye; ventral plates 130-137; subcaudals 78-86; apical pits very faint, small,absent on most scales (Schmidt 1925).

Detailed description in Malnate 1962: 261. 
Comment 
EtymologyNamed after American herpetologist Clifford Hillhouse Pope (1899-1974) who made important contributions to the herpetology of China. 
References
  • Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here
  • David, P., Bain, R. H., Nguyen Quang Truong, Orlov, N. L., Vogel, G., Vu Ngoc Thanh & T. Ziegler 2007. A new species of the natricine snake genus Amphiesma from the Indochinese Region (Squamata: Colubridae: Natricinae). Zootaxa 1462: 41-60 - get paper here
  • DAVID, PATRICK; GERNOT VOGEL, TRUONG QUANG NGUYEN, NIKOLAI L. ORLOV, OLIVIER S. G. PAUWELS, ALEXANDRE TEYNIÉ, THOMAS ZIEGLER 2021. A revision of the dark-bellied, stream-dwelling snakes of the genus Hebius (Reptilia: Squamata: Natricidae) with the description of a new species from China, Vietnam and Thailand. Zootaxa 4911: 1–61 - get paper here
  • Glass, Bryan P. 1946. A collection of reptiles from Hunan and Anhwei provinces, China. Copeia 1946 (4): 249-252 - get paper here
  • GUO, PENG; FEI ZHU, QIN LIU, LIANG ZHANG, JIAN X. LI, YU Y. HUANG & R. ALEXANDER PYRON 2014. A taxonomic revision of the Asian keelback snakes, genus Amphiesma (Serpentes: Colubridae: Natricinae), with description of a new species. Zootaxa 3873 (4): 425–440 - get paper here
  • Malnate, E.V. 1962. The relationships of five species of the Asiatic natricine snake genus Amphiesma. Proc. Acad. Nat. Sci. Philadelphia, 114, 251–299. - get paper here
  • Mell,R. 1931. List of Chinese snakes. Lingnan Sci. Jour., Canton, 8 [1929]: 199-219.
  • Nguyen, S.V., Ho, C.T. and Nguyen, T.Q. 2009. Herpetofauna of Vietnam. Chimaira, Frankfurt, 768 pp.
  • Resetar, A.R. & Voris, H. K. 1997. Herpetology at the Field Museum of Natural History, Chicago: the first one hundred years. in: Collection Building in Ichthyology and Herpetology, ASIH 1997, pp.495-506
  • Schmidt, KARL P. 1925. New reptiles and a new salamander from China. American Museum Novitates (157): 1-5. - get paper here
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp.
  • Wang, Kai; Jinlong Ren, Hongman Chen, Zhitong Lyu, Xianguang Guo Ke Jiang, Jinmin Chen, Jiatang Li, Peng Guo, Yingyong Wang, Jing Che 2020. The updated checklists of amphibians and reptiles of China. Biodiversity Science 28 (2): 189-218 - get paper here
  • Zhao, E.M. 2006. The snakes of China [in Chinese]. Hefei, China, Anhui Sience & Technology Publ. House, Vol. I, 372 pp., Vol. II (color plates), 280 pp.
  • Ziegler, T., and Quyet, L.K. 2006. A new natricine snake of the genus Amphiesma (Squamata: Colubridae: Natricinae) from the central Truong Son, Vietnam. Zootaxa 1225:39-56 - get paper here
 
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