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Heliobolus bivari MARQUES, CERÍACO, HEINICKE, CHEBOURI, CONRADIE, TOLLEY & BAUER, 2022

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Higher TaxaLacertidae, Eremiadinae, Sauria, Lacertoidea, Squamata (lizards)
Subspecies 
Common NamesE: Bivar’s bushveld lizard
Portuguese: lagartixa de Bivar 
SynonymHeliobolus bivari MARQUES, CERÍACO, HEINICKE, CHEBOURI, CONRADIE, TOLLEY & BAUER 2022 
DistributionAngola (southwestern xeric/desertic lowlands)

Type locality: Virulundo, Namibe Province, Angola (–16.2852°S, 12.9419°E), 718 m elevation  
Reproduction 
TypesHolotype: CAS 266287, adult female, collected by Luis M.P. Ceríaco, Suzana A. Bandeira and Ishan Agarwal on 2 December 2016.
Paratypes (n=16); CAS 266276-266286; PEM R17965, 17966, 21626; 21630, 24128, 24129 
DiagnosisDiagnosis: “A medium-sized lizard, identified to genus by the following combination of charac-ters: well-developed limbs, slender body, elongated snout, long tail, and a distinct collar on ventral region. Heliobolus bivari can be distinguished from other members of its genus by the following combination of characteristics: (1) stout medium-size body (mean SVL 54.6 mm); (2) long-tailed (mean 123.7 mm), tail more than twice the body length; (3) midbody scale rows 64–82 (mean: 71.5); (4) higher number of subdigital lamellae under the fourth toe (mean: 26.6); (5) parietals usually in contact, rarely separated; (6) cranial shields not ornamented and temporal shield smooth; (7) lateral dark marking through the ear to the posterior margin of the eye faded or totally absent; (8) absence of bright yellow dots ventrolaterally.”


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CommentDistribution: Possibly widespread to neighboring Namibia.

Diet: Expected to be similar to other Heliobolus species, therefore being oviparous and insectivorous.

Natural history: As commonly observed in juvenile H. lugubris individuals, it is assumed that juveniles of H. bivari mimic Anthia (Carabidae) ground beetles, both through their iconic white and yellow markings on a black body, and by adopting an “arched” position when walking, imitating the general shape of a beetle. 
EtymologyNamed after the Portuguese entomologist António Bivar de Sousa (Lisbon, 1946–), a researcher at the recently defunct Instituto de Investigação Ciemtífica Tropical (IICT), Lisboa, Portugal. Bivar de Sousa has had an important role in entomological research in Angola. Specific epithet is a patronym in the masculine genitive singular. 
References
  • Kwet, A. 2023. Liste der im Jahr 2022 neu beschriebenen Reptilien. Elaphe 2023 (3): 48-73
  • Marques MP, Ceríaco LMP, Heinicke MP, Chehouri RM, Conradie W, Tolley KA, Bauer AM 2022. The Angolan bushveld lizards, genus Heliobolus Fitzinger, 1843 (Squamata: Lacertidae): Integrative taxonomy and the description of two new species. Vertebrate Zoology 72: 745–769 - get paper here
 
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