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Hemidactylus kyaboboensis WAGNER, LEACHÉ & FUJITA, 2014

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common Names 
SynonymHemidactylus kyaboboensis WAGNER, LEACHÉ & FUJITA 2014: 3
Hemidactylus kyaboboensis LEACHÉ & FUJITA 2010 (nom. nud.) 
DistributionGhana

Type locality: Ghana, Volta Region, Togo Hills, Kyabobo National Park, Waterfall, 08.33019° N, 00.59411° E, 515 m  
Reproductionoviparous (not imputed, fide Zimin et al. 2022) 
TypesHolotype: MVZ 245291 (Fig. 2), adult male, collected by Adam D. Leaché, Raul Diaz and Matthew K. Fujita on 16 June 2004. Paratypes. MVZ 245292–245299, same data as the holotype. ZFMK 93689, subadult from Togo, Missahöhe [=Missahoe or Missahohé], near Kpalimé. ZFMK 19922, adult female from Faille de Bafilo [=fault proper near Bafilo (9.282399, 1.216621)], Togo. 
DiagnosisDiagnosis: A large species of Hemidactylus, with a max- imum SVL of 80 mm, and a maximum total length of 160 mm, a broad head (vs. slender in H. coalescens), and three enlarged internasal scales (vs. two in H. eniangii). Char- acteristic is the dorsal scalation of irregular arranged smooth and convex tubercle scales, each circumdated by a ring of small granular scales. In coloration, it is unique by having indistinct dark body crossbands, and more prominent whitish stripes and dots. This new species is distinct in coloration in detail to H. fasciatus by having a band on the side of the head as broad as the crossband on the neck, and reaching the lower tip of the ear hole (vs. not reaching the lower tip). It is distinct from H. coa- lescens by having the broad band on the side of the head (extension of the crossband on the neck) as broad as the band on the neck (vs. a crossband on the neck becoming a narrow stripe on the side of the head); by having the last indistinct body crossband in contact with the hindlimbs (vs. a distinct crossband not in contact); by having the first indistinct body crossband reaching the head skull (vs. a distinct first body crossband restricted to the neck). It is distinct from H. eniangii by having a band on the side of the head as broad as the crossband on the neck (vs. a cross- band becoming narrower on the side of the head).
 
CommentHabitat. At Togo Hills, specimens were collected in moist semi-deciduous rainforest, while the specimen at Missahöhe was found in an old stone building within moist semi-deciduous rainforest.

Relationships. This species includes all populations that cluster with those presented by Leaché & Fujita (2010) from the Togo Hills with strong support in the Bayesian species delimitation model. It is the sister taxon of H. fasciatus from the Guinean rainforest system in West Africa (Leaché & Fujita 2010; Leaché et al., 2014, Wagner et al. 2014).

Species delimitation: Douglas & Bouckaert 2022 applied a new species delimitation approach to this and other species and concluded that kyaboboensis is conspecific (synonymous) with H. fasciatus. 
EtymologyThis species is named after its type locality, the Kyabobo National Park, Togo Hills, Volta Region, Ghana. 
References
  • Bauer, Aaron M.; James F. Parham, Rafe M. Brown, Bryan L. Stuart, Lee Grismer, Theodore J. Papenfuss, Wolfgang Böhme, Jay M. Savage, Salvador Carranza, Jesse L. Grismer,, Philipp Wagner, Andreas Schmitz0, Natalia B. Ananjeva and Robert F. Inger 2010. Availability of new Bayesian-delimited gecko names and the importance of character-based species descriptions. Proc. R. Soc. B - doi:10.1098/rspb.2010.1330 - get paper here
  • Böhme, Wolfgang 2014. Herpetology in Bonn. Mertensiella 21. vi + 256 pp. - get paper here
  • Douglas, Jordan and Remco Bouckaert 2022. Quantitatively defining species boundaries with more efficiency and more biological realism. bioRxiv, February 16, 2022 - get paper here
  • Kwet, Axel 2015. Liste der im Jahr 2014 neu beschriebenen Reptilien. Terraria-Elaphe 2015 (3): 50-64 - get paper here
  • Leache, A. D., Oaks, J. R., Ofori‐Boateng, C. and Fujita, M. K. 2020. Comparative phylogeography of West African amphibians and reptiles. Evolution - get paper here
  • Rösler, Herbert 2015. Bemerkungen über einige Geckos der Zoologischen Staatssammlung München. Gekkota, Suppl. (2): 3-54
  • Segniagbeto, Gabriel Hoinsoude; Jean-François Trape, Komlan M. Afiademanyo, Mark-Oliver Rödel, Annemarie Ohler, Alain Dubois, Patrick David, Danny Meirte, Isabelle Adolé Glitho, Fabio Petrozzi, and Luca Luiselli 2015. Checklist of the lizards of Togo (West Africa), with comments on systematics, distribution, ecology, and conservation. Zoosystema 37 (2): 381-402 - get paper here
  • Wagner, P., Leaché, A. & Fujita, M.K. 2014. Description of four new West African forest geckos of the Hemidactylus fasciatus Gray, 1842 complex, revealed by coalescent species delimitation. Bonn zoological Bulletin 63 (1): 1–14 - get paper here
  • Zimin, A., Zimin, S. V., Shine, R., Avila, L., Bauer, A., Böhm, M., Brown, R., Barki, G., de Oliveira Caetano, G. H., Castro Herrera, F., Chapple, D. G., Chirio, L., Colli, G. R., Doan, T. M., Glaw, F., Grismer, L. L., Itescu, Y., Kraus, F., LeBreton 2022. A global analysis of viviparity in squamates highlights its prevalence in cold climates. Global Ecology and Biogeography, 00, 1–16 - get paper here
 
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