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Hemidactylus lanzai ŠMÍD, MAZUCH, NOVÁKOVÁ, MODRY, MALONZA, ABDIRAHMAN-ELMI, CARRANZA, AND MORAVEC, 2020

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common NamesE: Lanza’s gecko 
SynonymHemidactylus lanzai ŠMÍD, MAZUCH, NOVÁKOVÁ, MODRY, MALONZA, ABDIRAHMAN-ELMI, CARRANZA, AND MORAVEC 2020
Hemidactylus isolepis — BOULENGER 1896: 213
Hemidactylus isolepis — BOULENGER 1897: 277 (in part)
Hemidactylus isolepis — CALABRESI 1923: 151
Hemidactylus isolepis — LOVERIDGE 1923: 844
Hemidactylus isolepis — PARKER 1936: 601
Hemidactylus isolepis — LOVERIDGE 1947: 105 (in part)
Hemidactylus isolepis — DREWES 1972: 5
Hemidactylus isolepis — LANZA 1983: 203 (in part)
Hemidactylus isolepis — LANZA 1990: 415 (in part)
Hemidactylus isolepis — SPAWLS & ROTICH 1997: 64
Hemidactylus isolepis — SPAWLS et al. 2002: 87
Hemidactylus isolepis — LARGEN & SPAWLS 2006: 46 (in part)
Hemidactylus isolepis — LARGEN & SPAWLS 2010: 298 (in part)
Hemidactylus isolepis — RÖSLER 2015: 14 (in part)
Hemidactylus isolepis — SPAWLS et al. 2018: 89
Hemidactylus sp. 10 — ŠMÍD et al. 2013: 3
Hemidactylus sp. 10 — ŠMÍD et al. 2013: 81
Hemidactylus sp. 10 — GARCIA-PORTA et al. 2016: 3
Teratolepis isolepis — PARKER 1942: 35 
DistributionEthiopia

Type locality: Ethiopia, Southern Nations, Nationalities, and Peoples’ Region, 30 km SW of Arbore village, western bank of Chew Bahir Lake (4.844°N, 36.737°E, 628 m elevation)  
Reproduction 
TypesHolotype: NMP 6V 75857, adult male, 14 July 2014, collected by T. Mazuch and K. Kadlečková.
Paratypes: NMP6V 74448, subadult, Kenya, Marsabit County, Gas (3.065°N, 36.831°E, 460 m), September 2004, collected by D. Modrý. NMP6V 74449, NMP6V 74451/1–3, adult male (NMP6V 74451/1) and three sub-adults, Kenya, Marsabit County, 20 km E of Lake Turkana (2.532°N, 36.822°E, 760 m), September 2004, collected by D. Modrý. NMP6V 74450/1–3, one juvenile (74450/3) and two subadults, Kenya, Marsabit County, Kalacha, (3.136°N, 37.426°E, 375 m), September 2004, collected by D. Modrý. NMP6V 75656–75657, subadults, Kenya, Marsabit County, Korante Plain (1.977°N, 37.033°E, 920 m), September 2004, collected by D. Modrý. NMP6V 75743, adult female, Ethiopia, Southern Nations, Nationalities, and Peoples’ Region, 36 km E of Omorate on road to Turmi (4.739°N, 36.331°E, 565 m), 11 April 2016, collected by F. Kovařík. NMP75858, adult female, Ethiopia, Southern Nations, Nationalities, and Peoples’ Region, 23 km SE of Turmi (4.87°N, 36.648°E, 1133 m), 6 July 2013, collected by P. Novák and V. Trailin. MorphoBank picture numbers for the holotype, paratypes and 16 of the 36 referred specimens are provided in Supplemental Table S1. 
DiagnosisDiagnosis: A species of the African radiation of the arid clade of Hemidactylus (sensu ŠMÍD et al. 2013a) with the following morphological characteristics: (1) small size, SVL 29.6–41.0 mm in males, 34.1–42.4 mm in females; (2) moderately robust head, head depth 42–54% of head length, head width 68–81% of head length; (3) relatively short tail, tail length 103.8–116.2% of SVL; (4) uppermost nasals mostly in broad contact (exceptionally in point contact or separated); (5) large anterior postmentals in contact with the first and second infralabials, less frequently with the first infralabial alone; (6) seven to nine supralabials; (7) five to eight infralabials; (8) absence of enlarged dorsal tubercles; (9) dorsal scales homogeneous, smooth, and imbricate; (10) absence of tail segments or whorls; (11) subcaudals slightly enlarged or heterogeneous in size, imbricate; (12) 4–7 lamellae under the first finger, 6–9 lamellae under the fourth finger, 4–7 lamellae under the first toe, and 9–12 lamellae under the fourth toe; (13) four to eight precloacal pores in males; (14) presence of lateral postanal tubercles; (15) in life pinkish to pale brownish dorsally with irregular dark-brown spots and smaller white spots; the white spots are usually confined to several (up to five) adjoining scales, whereas the dark-brown spots are larger and tend to coalesce into transverse bands. Both dark and white dorsal spots present also on limbs, hands, and tail. A narrow dark-brown stripe runs from nostril across the eye to temporal area and is bordered from above by a pale stripe. A conspicuous dark brown triangular spot present on snout (Fig. 6 in ŠMÍD et al. 2019). Venter white or pinkish.


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CommentPublication date: while the paper gives the volume as 33 (2019), the fine print on page 45 says “Published on 13 March 2020”.

Distribution: See map in Smid et al. 2020: 32 (Fig. 3). 
EtymologyNamed after the late Prof. Benedetto Lanza (1924-2016), Italian herpetologist, in recognition of his crucial contribution to the knowledge of the herpetofauna of the Horn of Africa. 
References
  • Kirchhof S, Wasonga V, Mazuch T, Spawls S, Malonza KP. 2023. An annotated checklist of the herpetofauna of the Sibiloi National Park in northern Kenya based on field surveys. Amphibian & Reptile Conservation 17(1/2) [General Section]: 1–18 (e324) - get paper here
  • Spawls, Stephen; Tomáš Mazuch & Abubakr Mohammad 2023. Handbook of Amphibians and Reptiles of North-east Africa. Bloomsbury, 640 pp. - get paper here
  • Šmíd, Jiří, Mazuch, Tomáš, Nováková, Lucie, Modrý, David, Malonza, Patrick K., Abdirahman Elmi, Hassan Sh, Carranza, Salvador & Moravec, Jiří 2020. Phylogeny and Systematic Revision of the Gecko Genus Hemidactylus from the Horn of Africa (Squamata: Gekkonidae). Herpetological Monographs 33 (1): 26-47 [paper says 2019 on title page, but the fine print on page 45 says “Published on 13 March 2020] - get paper here
 
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