Hemidactylus granosus HEYDEN, 1827
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Higher Taxa | Gekkonidae, Gekkota, Sauria, Squamata (lizards: geckos) |
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Common Names | |
Synonym | Hemidactylus granosus HEYDEN 1827: 17 Hemidactylus verrucosus – RÜPPELL 1845: 300 (part.) Hemidactylus turcicus – BOETTGER 1893: 29 (part.) Hemidactylus turcicus – ANDERSON 1898: 80 (part.) Hemidactylus turcicus turcicus – LOVERIDGE 1947: 143 (part.) Hemidactylus turcicus turcicus – MERTENS & WERMUTH 1960: 79 (part.) Hemidactylus turcicus – SALVADOR 1981: 84 (part.) Hemidactylus turcicus – BAHA EL DIN 2006: 66 (part.) Hemidactylus turcicus turcicus – BAHA EL DIN 2005: 19 (part.) Hemidactylus turcicus turcicus – MERTENS 1967: 55 Hemidactylus sp. 1 – MORAVEC et al. 2011: 24 Hemidactylus sp. 1 – CARRANZA & ARNOLD 2012: 17 Hemidactylus sp. 1 – ŠMÍD et al. 2013: 3 Hemidactylus granosus — ŠMÍD et al. 2013 |
Distribution | Egypt (Sinai), SW Jordan, Saudi Arabia Type locality: “Ägypten, Arabien, und Abyssinien”. Restricted to “Arabia petraea” = Sinai, Egypt by MERTENS 1967. |
Reproduction | oviparous (manual and phylogenetic imputation, fide Zimin et al. 2022) |
Types | Lectotype SMF 8723 designated by Mertens (1967) |
Diagnosis | Diagnosis: Hemidactylus granosus is a member of the ‘Hemidactylus saba species group’ within the Arabian radiation of the Arid clade as evidenced by the mtDNA and nDNA analyses. The species has the following combination of molecular and morphological characters: (1) Uncorrected genetic distance from H. saba: 9.9–10.2% in 12S, 14.5–15.5% in cytb; from Hemidactylus sp. 4: 10.2–12.3% in 12S, 11.2–13.5% in cytb; (2) small size, SVL 39.0–53.2 mm in males, 40.6–53.3 mm in females; (3) rather elongated head, head length 24–28% of SVL, head width 68–86% of head length, head depth 33–47% of head length; (4) tail length 107–130% of SVL; (5) uppermost nasals separated by a small shield in 89% of specimens; (6) large anterior postmentals in wide mutual contact, and always in contact with the 1st and 2nd lower labial; (7) 9–11 upper labials; (8) 7–9 lower labials; (9) 14–15 longitudinal rows of enlarged, subtriangular, distinctly keeled dorsal tubercles; (10) 7–8 lamellae under the 1st toe and 10–13 under the 4th toe; (11) ca. 6–8 tail segments bearing 6 pointed tubercles; (12) 4–7 preanal pores in males forming a continuous row on the left and right side; (13) subcaudals enlarged; (14) in life, dorsum pale buff with dark brown spots tending to form transverse bands or X-shaped markings, dark horizontal stripe in prefrontal and temporal region, tail with ca. 10–13 dark brown transverse bands, venter white [from ŠMÍD et al. 2013]. |
Comment | Synonymy: Hemidactylus granosus HEYDEN 1827 has been resurrected from the synonymy of H. turcicus by SMID et al. 2013. Distribution: see map in ŠMÍD et al. 2016: Fig. 3, Pola et al. 2023: 4 (Fig. 1). |
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