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Hemidactylus hajarensis CARRANZA & ARNOLD, 2012

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common Names 
SynonymHemidactylus hajarensis CARRANZA & ARNOLD 2012
Hemidactylus persicus — ARNOLD, 1986: 419
Hemidactylus persicus — LEVITON et al. 1992: 38 (part.)
Hemidactylus persicus — VAN DER KOOIJ 2000: 112 (part.)
Hemidactylus persicus — CARRANZA & ARNOLD 2006: 536
Hemidactylus persicus — SINDACO & JEREMČENKO 2008: 115 (part.) 
DistributionN Oman (mountains from the Jebel Akhdar to the East), 22-1683 m elevation.

Type locality: Wadi Bani Khalid, 647 m, Eastern Hajar (North Oman), 22.61609’N 59.09371.  
Reproductionoviparous (not imputed, fide Zimin et al. 2022) 
TypesHolotype: BMNH 2008.714, male from Wadi Bani Khalid, 647 m, Eastern Hajar (North Oman), 22.61609’N 59.09371’E WGS84, collected in May 2011 by S. Carranza, E. Gómez-Díaz and F. Amat (MorphoBank M99903–M99917; Fig. 12D). Paratypes: ONHM3706, male, same collecting data as Holotype (MorphoBank M99885–M99893); IBES7335, male, same collecting data as Holotype (MorphoBank M99894–M99902; Fig. 11D); IBES7336, female, same collecting data as Holotype (MorphoBank M99969–M99976). 
DiagnosisDiagnosis A medium-sized Hemidactylus with a maximum recorded SVL of 66.9 mm; with a mean of 14.2 (13–15) longitudinal rows of enlarged dorsal tubercles at mid-body; adhesive pads fairly broad, in adults maximum width of pad on the fourth toe about 0.4–0.5 of its length; lamellae under the 1st toe of pes mean 8.0 (7–9); lamellae under the 4th toe of pes mean 12.1 (11–14); preanal pores mean 5.5 (4–6); expanded subcaudal scales extending proximally as far as the second or third whorl after the vent and starting just after the hemipenial bulge in males; dorsum grey-buff with irregular small spots; a dark stripe from the nostril, through the eye, on to cheek above ear and often on to neck; tail with small irregular dark blotches basally and numerous transverse dark bands more distally, the total number being around 17. Underside of tail pale but large subcaudals may be suffused with grey formed by dark chromatophores that increase in intensity distally; underside of toe pads also grey.
Hemidactylus hajarensis is generally similar to H. persicus in the number of its moderately-sized dorsal tubercles across mid-body, and large adhesive pads on toes but differs from it in its reduced number of preanal pores in males (mean 5.5, 4–6, compared with mean 9.2, 8–11). Hemidactylus hajarensis differs from H. luqueorum sp. nov. in its smaller size (SVL mean 54 mm, max. 66.9 mm, compared with mean. 76.8 mm, max. 88 mm), and in having fewer lamellae under the 1st toe of pes (mean 8.0, 7–9, compared with mean 10.3, 10–11), and under the 4th toe of pes (mean 12.1, 11–14, compared with mean 13.6, 13–14). 
CommentMorphoBank M94393–M94415 M94459–M94465 M94515–M94542 M94558–M94586 M94630–M94643 M94649–M94664 M94666–M94684 M94700–M94721 M99874–M99917 M99921–M99937 M99954–M99993 
EtymologyThe species epithet “hajarensis” is an adjective that refers to the mountain range where the species is found, the Hajar Mountains. 
References
  • Burriel-Carranza, B., Els, J., & Carranza, S. 2022. Reptiles & amphibians of the Hajar Mountains CSIC, Barcino Solucions Gràfiques, SL, Barcelona, - get paper here
  • Carranza S, Xipell M, Tarroso P, Gardner A, Arnold EN, Robinson MD, et al. 2018. Diversity, distribution and conservation of the terrestrial reptiles of Oman (Sauropsida, Squamata). PLoS One 13 (2): e0190389 - get paper here
  • Carranza, S., Arnold, E. Nicholas 2012. A review of the geckos of the genus Hemidactylus (Squamata: Gekkonidae) from Oman based on morphology, mitochondrial and nuclear data, with descriptions of eight new species. Zootaxa 3378: 1–95 - get paper here
  • Carranza, Salvador; Johannes Els; Bernat Burriel-Carranza 2021. A field guide to the reptiles of Oman. Madrid : Consejo Superior de Investigaciones Científicas, 223 pp. [review in HR 53 (3): 531] - get paper here
  • Gardner, A.S. 2013. The amphibians and reptiles of Oman and the UAE. Edition Chimaira, 480 pp.
  • Rösler, Herbert 2015. Bemerkungen über einige Geckos der Zoologischen Staatssammlung München. Gekkota, Suppl. (2): 3-54
  • Šmíd, Jiří; Salvador Carranza, Lukáš Kratochvíl, Václav Gvoždík, Abdul Karim Nasher, Jiří Moravec 2013. Out of Arabia: A Complex Biogeographic History of Multiple Vicariance and Dispersal Events in the Gecko Genus Hemidactylus (Reptilia: Gekkonidae). PLoS One 8(5): e64018 - get paper here
  • Zimin, A., Zimin, S. V., Shine, R., Avila, L., Bauer, A., Böhm, M., Brown, R., Barki, G., de Oliveira Caetano, G. H., Castro Herrera, F., Chapple, D. G., Chirio, L., Colli, G. R., Doan, T. M., Glaw, F., Grismer, L. L., Itescu, Y., Kraus, F., LeBreton 2022. A global analysis of viviparity in squamates highlights its prevalence in cold climates. Global Ecology and Biogeography, 00, 1–16 - get paper here
  • Zwanzig, Bernd-Michael & Wolfgang Grossmann 2013. Halbfingergeckos: Hemidactylus. Die verkannte Gattung. Terraria-Elaphe 2013 (2): 14-27 - get paper here
 
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