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Hemidactylus longicephalus BOCAGE, 1873

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common Names 
SynonymHemidactylus longicephalus BOCAGE 1873
Hemidactylus longiceps O’SHAUGHNESSY 1873 (nomen nudum)
Hemidactylus bocagii BOULENGER 1885: 125 (nom. nov. pro H. longiceps)
Hemidactylus mabouia var. molleri BEDRIAGA 1892
Hemidactylus hecqui BOULENGER 1901 (fide LOVERIDGE 1947)
Hemidactylus steindachneri TORNIER 1902 (fide LOVERIDGE 1947)
Hemidactylus heequi BOCAGE 1903: 63 (ex errore)
Hemidactylus longicephalus — SCHMIDT 1919: 445
Hemidactylus hecqui — WITTE 1933
Hemidactylus longicephalus — LOVERIDGE 1947
Hemidactylus longicephalus — KLUGE 1993
Hemidactylus longicephalus — RÖSLER 2000: 87
Hemidactylus longicephalus — CERIACO et al. 2014
Hemidactylus longicephalus — CERIACO et al. 2020: 33 
DistributionAngola (Bengo, Kwanza Sul, Kwanza Norte, Luanda, Benguela), W Democratic Republic of the Congo (Zaire), Cameroon; São Tomé, Principé, Central African Republic, Namibia, Equatorial Guinea

Type locality: Capangombe and Catumbella, Angola.
Neotype locality: Kawa Camp, Kissama National Park (-9.18303°, 13.37063°, 136 m), Luanda Province, Republic of Angola  
Reproductionoviparous 
TypesNeotype: CAS 263575 (field number MCZ A-36523; Fig. 15; Table 4), adult male, collected by Luis M.P. Ceríaco, Mariana P. Marques, Philip Pastor and John Cavagnaro on 13 June 2016, designated by Ceríaco et al. 2020: 38. Original holotype lost, was MB (Museu Bocage) (probably lost in fire 1978).
Neotype: CAS 219293 (field number CAS-RCD 14215; Fig. 18; Table 4) adult male collected in Coast road south of Shipwreck Cove (0.38739° N, 6.59492° E; 28 m), Lobata District, São Tomé Island, Republic of São Tomé and Príncipe, collected by R.E. Stoelting on 3 May 2001, designated by Ceríaco et al. 2020: 42 [molleri] 
DiagnosisDiagnosis: A medium sized Hemidactylus, maximum snout-vent length 54.8 mm (Fig. 14). Dorsal pholidosis heterogeneous, with 16–18 irregularly arranged longitudinal rows of subtrihedral, striated, moderately keeled tubercles. Two well-developed pairs of postmentals, the inner pair slightly longer than the outer pair, about two size of the mental, and in broad contact behind the mental. ventrolateral folds distinct, about 30–34 scale rows across venter. Six or seven divided scansors beneath first digit of both manus and pes, seven or eight beneath fourth digit of manus, nine or ten (rarely eight) beneath the fourth digit of pes. Males with 4–11 precloacal-femoral pores in a continuous series. Original tail slightly cyclotetragonal, strongly spinose; dorsal scales on tail slightly larger than dorsals of body, striated, imbricate, with a longitudinal series of two enlarged, strongly keeled, striated, pointed tubercles on either side of the median dorsal furrow. Subcaudal scales small and imbricate, about one fifth of tail width. Dorsum greyish with a series of sometimes incomplete dark-brown transverse markings from occiput to sacrum, sometimes bordered by a thin cream-colored stripe extending from behind the eyes to the insertion of the forelimbs; tail with distinct alternating light and dark bands. 
CommentSynonymy after KLUGE 1993. See also Ceríaco et al. 2020: 33.

Distribution: Not listed for Tanzania by BROADLEY & HOWELL 1991. Not in East Africa (i.e. Kenya, Tanzania, Uganda, Rwanda, Burundi fide S. Spawls, pers. comm., 28 June 2018). For a map with localities in Equatorial Guinea see SÁNCHEZ-VIALAS et al. 2022. 
References
  • Bedriaga, J. von. 1892. Notes sur les amphibiens et reptiles recueillis par M. Adolphe F. Moller aux Iles de la Guinée. O. Instituto; Revista Cientifica e Litteraria, Coimbra, 39: 498–507; 642–648; 736–742; 814–820; 901–907 (reprint pagination: 1-48).
  • Bocage, Barboza du 1903. G.A. Boulenger - Batraciens nouveaux et Reptiles nouveaux. J. Sci. math. phys. nat. Lisboa: 7 (2): 62-64
  • Bocage, J.V.B. 1873. Melanges erpetologiques. II. Sur quelques reptiles et batraciens nouveaux, rares ou peu connus d‘Afrique occidentale. Jorn. Sci. Math. Phys. Nat. Lisboa 4: 209-227 - get paper here
  • Boulenger, G. A. 1901. Materiaux pour la faune du Congo. Batraciens et reptiles nouveaux. Ann. Mus. Congo Belge, Zool. (sect. C. ser. 1) 2: 7-14.
  • Boulenger, G.A. 1885. Catalogue of the lizards in the British Museum (Nat. Hist.) I. Geckonidae, Eublepharidae, Uroplatidae, Pygopodidae, Agamidae. London: 450 pp. - get paper here
  • Ceríaco, L. M. P., Marques, M. P., de Sousa, A. C. A., Veríssimo, J., Beja, P., & Ferreira, S. 2023. Illustrated keys and a DNA barcode reference library of the amphibians and terrestrial reptiles (Amphibia, Reptilia) of São Tomé and Príncipe (Gulf of Guinea, West Africa). ZooKeys, 1168, 41-75
  • Ceríaco, Luis M. P., Aaron M. Bauer, David C. Blackburn and Ana C. F. C. Lavres. 2014. The herpetofauna of the Capanda Dam Region, Malanje, Angola. Herpetological Review 45 (4): 667-674 - get paper here
  • CERÍACO, LUIS M. P.; ISHAN AGARWAL, MARIANA P. MARQUES, AARON M. BAUER 2020. A review of the genus Hemidactylus Goldfuss, 1820 (Squamata: Gekkonidae) from Angola, with the description of two new species. Zootaxa 4746 (1): 1-71. - get paper here
  • CERÍACO, LUIS M. P.; ISHAN AGARWAL, MARIANA P. MARQUES, AARON M. BAUER 2020. A correction to a recent review of the genus Hemidactylus Goldfuss, 1820 (Squamata: Gekkonidae) from Angola, with the description of two additional species. Zootaxa 4861 (1): 92-106 - get paper here
  • Chirio, L. & Lebreton, M. 2007. Atlas des reptiles du Cameroun. MNHN, IRD, Paris 688 pp.
  • Chirio, Laurent and Ivan Ineich 2006. Biogeography of the reptiles of the Central African Republic. African Journal of Herpetology 55(1):23-59. - get paper here
  • Ernst R, Lautenschläger T, Branquima MF, Hölting M 2020. At the edge of extinction: a first herpetological assessment of the proposed Serra do Pingano Rainforest National Park in Uíge Province, northern Angola. Zoosystematics and Evolution 96(1): 237-262 - get paper here
  • Herrmann, H.-W.; W.R. Branch 2013. Fifty years of herpetological research in the Namib Desert and Namibia with an updated and annotated species checklist. Journal of Arid Environments 93: 94–115 - get paper here
  • Lobón-Rovira, J., Conradie, W., Buckley Iglesias, D., Ernst, R., Veríssimo, L., Baptista, N., & Pinto, P. V. 2021. Between sand, rocks and branches: an integrative taxonomic revision of Angolan Hemidactylus Goldfuss, 1820, with description of four new species. Vertebrate Zoology 71: 465–501; DOI 10.3897/vz.73.e64781 - get paper here
  • Loveridge, A. 1947. Revision of the African lizards of the family Gekkondiae. Bull. Mus. Comp. Zool. Harvard 98: 1-469 - get paper here
  • Marques, Mariana P.; Luis M. P. Ceríaco , David C. Blackburn , and Aaron M. Bauer 2018. Diversity and Distribution of the Amphibians and Terrestrial Reptiles of Angola -- Atlas of Historical and Bibliographic Records (1840–2017). Proc. Cal. Acad. Sci. (Ser. 4) 65: 1-501 (Supplement II)
  • Miller, Elizabeth C.; Anna B. Sellas & Robert C. Drewes 2012. A new species of Hemidactylus (Squamata: Gekkonidae) from Príncipe Island, Gulf of Guinea, West Africa with comments on the African-Atlantic clade of Hemidactylus geckos. African Journal of Herpetology 61 (1): 40-57
  • O’Shaughnessy, A. W. E. 1873. Reptilia. Zoological Record, 10:75—98
  • Pollo, César J. 2017. Guide of Amphibians and Reptiles of São Tomé and Príncipe. Smashwords / César J. Pollo, 156 pp. - get paper here
  • Rösler, H. 2000. Kommentierte Liste der rezent, subrezent und fossil bekannten Geckotaxa (Reptilia: Gekkonomorpha). Gekkota 2: 28-153
  • Rösler, Herbert 2015. Bemerkungen über einige Geckos der Zoologischen Staatssammlung München. Gekkota, Suppl. (2): 3-54
  • SÁNCHEZ-VIALAS, A., CALVO-REVUELTA, M. & DE LA RIVA, I. 2022. Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197 - get paper here
  • SANTOS, B. S. .,MARQUES, M. P. ., BAUER, A. M. ., & CERÍACO, L. M. P. . 2021. Herpetological results of Francisco Newton’s Zoological Expedition to Angola (1903–1906): a taxonomic revision and new records of a forgotten collection. Zootaxa 5028 (1): 1–80 - get paper here
  • Schmidt, Karl Patterson 1919. Contributions to the Herpetology of the Belgian Congo based on the Collection of the American Congo Expedition, 1909-1915. Part I: turtles, crocodiles, lizards, and chamaeleons. Bull. Amer. Mus. Nat. Hist. 39 (2): 385-624 - get paper here
  • Tornier, G. 1902. Die Crocodile, Schildkröten und Eidechsen in Kamerun. Zool. Jahrb., Abt. Syst., 15 (6): 663-677 - get paper here
  • Witte, G. F. de 1933. Reptiles récoltés au Conge Belge par le Dr. H. Schouteden et par M. G.-F. Witte. Ann. Mus. Conge belge Zool. Ser. 1 Tome III: 53-100.
  • Witte, G. F. de 1953. Reptiles. Exploration du Parc National de l'Upemba. Mission G. F. de Witte en collaboration avec W. Adam, A. Janssens, L. Van Meel et R. Verheyen (1946–1949). Institut des Parcs Nationaux du Congo Belge. Brussels, vol. 6, 322 pp. - get paper here
 
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