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Hemidactylus mandebensis ŠMÍD, MORAVEC, KRATOCHVIL, NASHER, MAZUCH, GVOŽDÍK & CARRANZA, 2015

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common Names 
SynonymHemidactylus mandebensis SMID, MORAVEC, KRATOCHVIL, NASHER, MAZUCH, GVOZDIK & CARRANZA 2015
Hemidactylus sp. 5 — MORAVEC et al. 2011
Hemidactylus sp. 6 — SMID et al. 2013 
DistributionYemen

Type locality: Yemen, Ta’izz governorate, 3 km S of Najd an Nashamah, 13.358° N, 43.957° E, 1182 m elevation  
Reproductionoviparous (manual imputation, fide Zimin et al. 2022) 
TypesHolotype: NMP P6V 74836/2, adult male (MorphoBank M329385-M329406), 26. X. 2007, collected by L. Kratochvıl. Paratypes. NMP6V 74836/1, adult female (MorphoBank M329505-M329516), same data as holotype; NMP6V 74970, subadult female (MorphoBank M329407- M329416), Yemen, Ta’izz governorate, Jabal Sabir, 13.528°N, 43.952°E, 1253 m elevation, 26. X. 2005, collected by L. Kratochvıl 
DiagnosisDiagnosis: A small-sized species of the H. robustus spe- cies group within the arid clade of Hemidactylus characterized by: (1) small size with maximum recorded SVL 41.5 mm in the only known male and 39.1 mm in females; (2) 78 infralabials and 9 supralabials; (3) infraorbital, parietal and temporal region covered with numerous round unkeeled tubercles; (4) large anterior postmentals in wide mutual contact; (5) dorsum with 1214 longitudinal rows of round, enlarged, keeled, and posteriorly slightly pointed tubercles; (6) 6 preanal pores in males (only one male known); (7) 56 lamellae under the 1st toe; (8) 89 lamellae under the 4th toe; (9) enlarged subcaudals; (10) in life having dark brown irregular pattern on greyish body colour, two prominent dark stripes from eye to the shoulder region and bright white dorsal tubercles. 
CommentDistribution: see map in ŠMÍD et al. 2016: Fig. 3. 
EtymologyThe species epithet is an adjective that refers to the Bab-el-Mandeb strait, the southern gate to the Red Sea in the vicinity of which the species occurs. 
References
  • MORAVEC, JIŘÍ; LUKÁŠ KRATOCHVÍL, ZUAIR S. AMR, DAVID JANDZIK, JIŘÍ ŠMÍD, & VÁCLAV GVOŽDÍK 2011. High genetic differentiation within the Hemidactylus turcicus complex (Reptilia: Gekkonidae) in the Levant, with comments on the phylogeny and systematics of the genus. Zootaxa 2894: 21–38
  • Šmíd J, Moravec J, Kratochvíl L, Gvoždík V, Nasher AK, Busais SM, Wilms T, Shobrak MY, Carranza S 2013. Two newly recognized species of Hemidactylus (Squamata, Gekkonidae) from the Arabian Peninsula and Sinai, Egypt. ZooKeys 355: 79–107 - get paper here
  • Šmíd, Jiří et al. 2015. Multilocus phylogeny and taxonomic revision of the Hemidactylus robustus species group (Reptilia, Gekkonidae) with descriptions of three new species from Yemen and Ethiopia. Systematics and Biodiversity, DOI: 10.1080/14772000.2014.996264 - get paper here
  • Šmíd, Jiří; Marek Uvizl, Mohammed Shobrak, Salem Busais, Al Faqih Ali Salim, Raed Hamoud M. AlGethami, Abdulaziz Raqi AlGethami, Abdulkarim Saleh K. Alanazi, Saad Dasman Alsubaie, Michail Rovatsos, Lucie Nováková, Tomáš Mazuch & Salvador Carranza 2022. Diversification of Hemidactylus geckos (Squamata: Gekkonidae) in coastal plains and islands of southwestern Arabia with descriptions and complete mitochondrial genomes of two endemic species to Saudi Arabia. Organisms Diversity & Evolution, 1-23 - get paper here
  • Šmíd, Jiří; Salvador Carranza, Lukáš Kratochvíl, Václav Gvoždík, Abdul Karim Nasher, Jiří Moravec 2013. Out of Arabia: A Complex Biogeographic History of Multiple Vicariance and Dispersal Events in the Gecko Genus Hemidactylus (Reptilia: Gekkonidae). PLoS One 8(5): e64018 - get paper here
  • Šmíd, Jiří; Mohammed Shobrak; Thomas Wilms; Ulrich Joger & Salvador Carranza 2016. Endemic diversification in the mountains: genetic, morphological, and geographical differentiation of the Hemidactylus geckos in southwestern Arabia. Org Divers Evol, DOI 10.1007/s13127-016-0293-3 - get paper here
  • Zimin, A., Zimin, S. V., Shine, R., Avila, L., Bauer, A., Böhm, M., Brown, R., Barki, G., de Oliveira Caetano, G. H., Castro Herrera, F., Chapple, D. G., Chirio, L., Colli, G. R., Doan, T. M., Glaw, F., Grismer, L. L., Itescu, Y., Kraus, F., LeBreton 2022. A global analysis of viviparity in squamates highlights its prevalence in cold climates. Global Ecology and Biogeography, 00, 1–16 - get paper here
 
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