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Hemidactylus gramineus CERÍACO, BAUER, KUSAMBA, AGARWAL & GREENBAUM, 2021

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common NamesFrench: ‘Gecko de l’herbe
E: grass-dwelling tropical gecko 
SynonymHemidactylus gramineus CERÍACO, BAUER, KUSAMBA, AGARWAL & GREENBAUM 2021
Hemidactylus cf. nzingae — CERÍACO et al. 2020: 14
Hemidactylus sp. — CERÍACO et al. 2020: 19 
DistributionDemocratic Republic of Congo (Katanga; Kinshasa: Bombo- Lumene Reserve)

Type locality: bank of Lumene River, Bombo-Lumene Reserve (-4.420278, 16.046878, 548 m), Kinshasa Province, DRC.  
Reproduction 
TypesHolotype. UTEP 22208 (field number ELI 1645; Figs. 1, 2), adult male, collected by Chifundera Kusamba and Wandege M. Muninga on 7 June 2013.
Paratype. UTEP 22209 (field number ELI 1674), adult male collected in Bombo-Lumene Reserve (-4.418438, 16.044208, 555 m), Kinshasa Province, Democratic Republic of the Congo, collected by Chifundera Kusamba, Wandege M. Muninga, Mwenebatu M. Aristote, and Eli Greenbaum on 7 June 2013. 
DiagnosisDiagnosis: A small-sized Hemidactylus, maximum SVL 39.5 mm, with a short, blunt snout (Fig. 1). Dorsal pholidosis heterogeneous, with 12–14 irregularly arranged longitudinal rows of subtrihedral, striated, strongly keeled tubercles at midbody. Two well-developed pairs of postmentals, the inner pair longer than the outer pair, about half the size of the mental, and in contact behind the mental. Ventrolateral folds distinct, 23– 25 scale rows across midventer. Five divided scansors beneath first digit of both manus and pes, seven beneath fourth digit of manus, seven beneath the fourth digit of pes. Males with a continuous series of eight precloacal pores. Body dorsum grayish brown, bordered by irregular, squiggly, dark brown markings that are interrupted by pairs of light gray spots, bordered by a cream/tan band that passes from the tip of the snout through the eye to the upper portion of the flanks and onto the tail, with numerous whitish tubercles on the flanks (CERÍACO et al. 2021).


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Comment 
EtymologyThe specific epithet ‘‘gramineus’’ refers to the grassy type of habitat from which the species was collected in the DRC and the Republic of the Congo (Fig. 5) and is applied here as an adjective in the genitive singular. 
References
  • Ceríaco, L. M., Bauer, A. M., Kusamba, C., Agarwal, I., & Greenbaum, E. 2021. A New Species of Ground-Dwelling Hemidactylus (Squamata: Gekkonidae) from Southwestern Democratic Republic of the Congo. Journal of Herpetology 55 (2): 105-111 - get paper here
  • CERÍACO, LUIS M. P.; ISHAN AGARWAL, MARIANA P. MARQUES, AARON M. BAUER 2020. A review of the genus Hemidactylus Goldfuss, 1820 (Squamata: Gekkonidae) from Angola, with the description of two new species. Zootaxa 4746 (1): 1-71. - get paper here
  • Lobón-Rovira, J., Conradie, W., Buckley Iglesias, D., Ernst, R., Veríssimo, L., Baptista, N., & Pinto, P. V. 2021. Between sand, rocks and branches: an integrative taxonomic revision of Angolan Hemidactylus Goldfuss, 1820, with description of four new species. Vertebrate Zoology 71: 465–501; DOI 10.3897/vz.73.e64781 - get paper here
 
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