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Hemiphyllodactylus bintik GRISMER, WOOD, ANUAR, QUAH, MUIN, ONN, SUMARLI & LOREDO, 2015

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common NamesE: SPOTTED DWARF GECKO
Malay: CICAK KERDIL BERINTIK 
SynonymHemiphyllodactylus bintik GRISMER, WOOD, ANUAR, QUAH, MUIN, ONN, SUMARLI & LOREDO 2015 
DistributionWest Malaysia (Terengganu)

Type locality: at 800 m elevation on Gunung Tebu, Terengganu, Peninsular Malaysia (05°36.11′N, 102°36.19′S).  
Reproductionoviparous (not imputed, fide Zimin et al. 2022) 
TypesHolotype: LSUHC 11216, adult male collected by Chan Kin Onn on 2 July 2013 
DiagnosisDiagnosis: Hemiphyllodactylus bintik sp. nov. can be separated from all other species of Hemiphyllodactylus by the unique combination of: a maximum SVL of 36.6 mm; seven chin scales extending transversely from unions of second and third infralabials and posterior margin of mental; enlarged postmental scales; five circumnasal scales; three scales between supranasals (= postrostrals); 11 supralabials; 12 infralabials; 17 longitudinally arranged dorsal scales at midbody contained within one eye diameter; seven longitudinally arranged ventral scales at midbody contained within one eye diameter; lamellar formula on hand 2443; lamellar formula on foot 3444; dorsal body pattern consisting of dark, squarish, paravertebral blotches; postsacral mark cream-coloured, lacking anteriorly projecting arms; and caecum and oviducts unpigmented. These characters and potentially diagnostic morphometric characters are scored across all species in Table 4 in Grismer et al. 2015.
 
CommentSympatry: H. tehtarik is found in the vicinity of H. bintik

Abundance: only known from the type specimen (Meiri et al. 2017). 
EtymologyThe specific epithet ‘bintik’ is a Malaysian word for ‘spot’, and refers to the spotted dorsal pattern of this species. 
References
  • COBOS, ANTHONY; L. LEE GRISMER, PERRY L. WOOD, JR., EVAN S. H. QUAH, SHAHRUL ANUAR, MOHD ABDUL MUIN 2016. Phylogenetic relationships of geckos of the Hemiphyllodactylus harterti group, a new species from Penang Island, Peninsular Malaysia, and a likely case of true cryptic speciation. Zootaxa 4107 (3): 367-380 - get paper here
  • GRISMER, L. LEE; & EVAN S. H. QUAH 2019. An updated and annotated checklist of the lizards of Peninsular Malaysia, Singapore, and their adjacent archipelagos. Zootaxa 4545 (2): 230–248 - get paper here
  • Grismer, L. Lee; Perry L. Wood Jr, Shahrul Anuar, Evan S. H. Quah, Mohd Abdul Muin, Chan Kin Onn, Alexandra X. Sumarli and Ariel I. Loredo 2015. Repeated evolution of sympatric, palaeoendemic species in closely related, co-distributed lineages of Hemiphyllodactylus Bleeker, 1860 (Squamata: Gekkonidae) across a sky-island archipelago in Peninsular Malaysia. Zoological Journal of the Linnean Society 174 (4): 859–876; DOI: 10.1111/zoj.12254 - get paper here
  • Meiri, Shai; Aaron M. Bauer, Allen Allison, Fernando Castro-Herrera, Laurent Chirio, Guarino Colli, Indraneil Das, Tiffany M. Doan, Frank Glaw, Lee L. Grismer, Marinus Hoogmoed, Fred Kraus, Matthew LeBreton, Danny Meirte, Zoltán T. Nagy, Cristiano d 2017. Extinct, obscure or imaginary: the lizard species with the smallest ranges. Diversity and Distributions - get paper here
  • NGUYEN, TRUONG QUANG; QUYEN HANH DO, HANH THI NGO, ANH VAN PHAM,<br>CUONG THE PHAM, MINH DUC LE, & THOMAS ZIEGLER 2020. Two new species of Hemiphyllodactylus (Squamata: Gekkonidae) from Hoa Binh Province, Vietnam. Zootaxa 4801 (3): 513–536 - get paper here
  • SUKPRASERT, APISAK; SURAPOP SUTTHIWISES, VIRAYUTH LAUHACHINDA, WUT TAKSINTUM 2018. Two new species of Hemiphyllodactylus Bleeker (Squamata: Gekkonidae) from Thailand. Zootaxa 4369 (3): 363–376 - get paper here
  • Zimin, A., Zimin, S. V., Shine, R., Avila, L., Bauer, A., Böhm, M., Brown, R., Barki, G., de Oliveira Caetano, G. H., Castro Herrera, F., Chapple, D. G., Chirio, L., Colli, G. R., Doan, T. M., Glaw, F., Grismer, L. L., Itescu, Y., Kraus, F., LeBreton 2022. A global analysis of viviparity in squamates highlights its prevalence in cold climates. Global Ecology and Biogeography, 00, 1–16 - get paper here
 
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