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Hemiphyllodactylus banaensis NGO VAN TRI, GRISMER, THAI & WOOD, 2014

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Common NamesE: Ba Na Dwarf Gecko 
SynonymHemiphyllodactylus banaensis NGO VAN TRI, GRISMER, THAI & WOOD 2014
Hemiphyllodactylus banaensis — LUU et al. 2023 
DistributionVietnam (HoaVang District)

Type locality: Ba Na summit of Ba Na–Nui Chua Nature Reserve, HoaVang District, Da Nang City, Vietnam (15°59’30’’N, 107°59’50’’E; 1300 m elevation)  
Reproductionoviparous (not imputed, fide Zimin et al. 2022) 
TypesHolotype: ITBCZ 2465, adult male, collected on 2 February 2013 by Pham Hong Thai at 21:45hrs; Paratypes. Eight additional specimens collected by Pham Hong Thai very near the type locality but on different days: ITBCZ 2450 on 19 August 2012, ITBCZ 2461–2464 on 23 September 2012, ITBCZ 2466–67 and ITBCZ 2469 have the same data as the holotype. ITBCZ 2468 was collected by Ngo Van Tri on 2 February 2013. 
DiagnosisDiagnosis: Hemiphyllodactylus banaensis sp. nov. can be distinguished from all other species of Hemiphyllodactylus by having the unique combination of a maximum SVL of 48.2 mm in males and 51.0 mm in females; seven chin scales extending transversely from the union of the first and second infralabials and posterior margin of mental; enlarged postmental scales; three circumnasal scales; 4–11 scales between supranasals; 9–12 supralabials; 9–11 infralabials;18–20 longitudinally arranged dorsal scales at midbody contained within one eye diameter; 9–12 longitudinally arranged ventral scales at midbody contained within one eye diameter; lamellar formula on hand 3 (II), 4–5 (III), 4–5 (IV), 4 (V); lamellar formula on foot 4 (II), 5 (III), 5 (IV), 5 (V); 20–21 precloacal and femoral pore-bearing scales contiguous in males and 0–20 contiguous pore-bearing precloacal scales in females; dorsal pattern on body composed of transverse blotches and two whitish stripes across shoulder extending to sacrum; postsacral mark whitish brown and bearing anteriorly projecting arms; and caecum and oviducts unpigmented. These characters and potentially diagnostic morphometric characters are scored across all species of Hemiphyllodactylus in Table 1 (TRI et al. 2014). 
CommentAbundance: only known from its original description (Meiri et al. 2017). 
EtymologyThis specific epithet is derived from the name of Ba Na Mountain and is in reference to the type locality. 
  • GRISMER, L. LEE; AWAL RIYANTO, DJOKO T. ISKANDAR & JIMMY A. MCGUIRE 2014. A new species of Hemiphyllodactylus Bleeker, 1860 (Squamata: Gekkonidae) from Pulau Enggano, southwestern Sumatra, Indonesia. Zootaxa 3821 (4): 485–495 - get paper here
  • Grismer, L. Lee; Perry L. Wood Jr, Shahrul Anuar, Evan S. H. Quah, Mohd Abdul Muin, Chan Kin Onn, Alexandra X. Sumarli and Ariel I. Loredo 2015. Repeated evolution of sympatric, palaeoendemic species in closely related, co-distributed lineages of Hemiphyllodactylus Bleeker, 1860 (Squamata: Gekkonidae) across a sky-island archipelago in Peninsular Malaysia. Zoological Journal of the Linnean Society 174 (4): 859–876; DOI: 10.1111/zoj.12254 - get paper here
  • GUO, WEIBO; KAIYA ZHOU, JIE YAN & PENG LI 2015. A new species of Hemiphyllodactylus Bleeker, 1860 (Squamata: Gekkonidae) from western Yunnan, China. Zootaxa 3974 (3): 377–390 - get paper here
  • Luu VQ, Nguyen TH, Do QH, Pham CT, Hoang TT, Nguyen TQ, Le MD, Ziegler T, Grismer JL, Grismer LL 2023. A new species of Hemiphyllodactylus (Squamata, Gekkonidae) from Ha Giang Province, Vietnam. ZooKeys 1167: 353-382 - get paper here
  • Meiri, Shai; Aaron M. Bauer, Allen Allison, Fernando Castro-Herrera, Laurent Chirio, Guarino Colli, Indraneil Das, Tiffany M. Doan, Frank Glaw, Lee L. Grismer, Marinus Hoogmoed, Fred Kraus, Matthew LeBreton, Danny Meirte, Zoltán T. Nagy, Cristiano d 2017. Extinct, obscure or imaginary: the lizard species with the smallest ranges. Diversity and Distributions - get paper here
  • NGO VAN TRI; L. LEE GRISMER, PHAM HONG THAI & P. L. WOOD, JR. 2014. A new species of Hemiphyllodactylus Bleeker, 1860 (Squamata: Gekkonidae) from Ba Na–Nui Chua Nature Reserve, Central Vietnam. Zootaxa 3760 (4): 539–552 - get paper here
  • NGUYEN, TRUONG QUANG; QUYEN HANH DO, HANH THI NGO, ANH VAN PHAM,<br />CUONG THE PHAM, MINH DUC LE, & THOMAS ZIEGLER 2020. Two new species of Hemiphyllodactylus (Squamata: Gekkonidae) from Hoa Binh Province, Vietnam. Zootaxa 4801 (3): 513–536 - get paper here
  • SUKPRASERT, APISAK; SURAPOP SUTTHIWISES, VIRAYUTH LAUHACHINDA, WUT TAKSINTUM 2018. Two new species of Hemiphyllodactylus Bleeker (Squamata: Gekkonidae) from Thailand. Zootaxa 4369 (3): 363–376 - get paper here
  • SUNG, YIK-HEI; WING-HO LEE, HO-NAM NG, YANJIE ZHANG, JIAN-HUAN YANG 2018. A new species of Hemiphyllodactylus (Squamata: Gekkonidae) from Hong Kong. Zootaxa 4392 (2): 361–373 - get paper here
  • Zimin, A., Zimin, S. V., Shine, R., Avila, L., Bauer, A., Böhm, M., Brown, R., Barki, G., de Oliveira Caetano, G. H., Castro Herrera, F., Chapple, D. G., Chirio, L., Colli, G. R., Doan, T. M., Glaw, F., Grismer, L. L., Itescu, Y., Kraus, F., LeBreton 2022. A global analysis of viviparity in squamates highlights its prevalence in cold climates. Global Ecology and Biogeography, 00, 1–16 - get paper here
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