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Hemiphyllodactylus indosobrinus ELIADES, PHIMMACHAK, SIVONGXAY, SILER & STUART, 2019

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common Names 
SynonymHemiphyllodactylus indosobrinus ELIADES, PHIMMACHAK, SIVONGXAY, SILER & STUART 2019: 138
Hemiphyllodactylus sp. nov. 8 — GRISMER et al. 2015:863
Hemiphyllodactylus sp. nov. 9 — GRISMER et al. 2013:852;
Hemiphyllodactylus sp. nov. 9 — GRISMER et al. 2014:490
Hemiphyllodactylus sp. nov. 9 — GRISMER et al. 2014:541
Hemiphyllodactylus sp. nov. 9 — NGUYEN et al. 2014:46
Hemiphyllodactylus sp. nov. 9 — YAN et al. 2016:544
Hemiphyllodactylus sp. nov. 9 — GRISMER et al. 2017:6
Hemiphyllodactylus sp. nov. 9 — SUKPRASERT et al. 2018:365
Hemiphyllodactylus cf. yunnanensis — NGUYEN et al. 2013:91 
DistributionLaos (Champasak Province)

Type locality: Bolaven Plateau, Dong Hua Sao National Protected Area, Pakxong District, Champasak Province, Laos, 15.07694°N, 106.13750°E, WGS84 (Fig. 1A in Eliades et al. 2019), ca. 1,000 m elev  
Reproductionoviparous (manual imputation, fide Zimin et al. 2022) 
TypesHolotype. FMNH 258695 (field number HKV 63933), adult male, collected 13 September 1999 by BLS, Harold F. Heatwole, and Bee Thaovanseng. 
DiagnosisDiagnosis. Hemiphyllodactylus indosobrinus sp. nov. can be distinguished from all congeners by having the following combination of characters: supralabials 15; infralabials 12; precloacofemoral pores 18 in males (females unknown); subdigital lamellae on Fingers II–V 4-5-5-4; total lamellae on hand 18; subdigital lamellae on Toes II– V 4-5-5-5; total lamellae on foot 19; dorsal scales across midbody within one orbital diameter 30; ventral scales across midbody within one orbital diameter 11; cloacal spurs two; chin scales eight; and internasal scales three.

Comparisons. Hemiphyllodactylus indosobrinus sp. nov. can be differentiated from all other congeners in mainland Southeast Asia and southern China by a combination of the following characters: chin scales eight (versus ≥10 in H. aurantiacus and ≤6 in H. linnwayensis Grismer, Wood, Thura, Zin, Quah, Murdoch, Grismer, Li, Kyaw, & Lwin, 2017); postmentals distinctly enlarged (versus not enlarged in H. aurantiacus); circumnasal scales three (versus ≥4 in H. flaviventris, H. khlonglanensis Sukprasert, Sutthiwises, Lauhachinda, & Taksintum, 2018, H. kiziriani, and H. linnwayensis); supralabial scales 15 [versus ≤13 in H. aurantiacus, H. banaensis, H. changningensis Guo, Zhou, Yan, & Li, 2015, H. chiangmaiensis Grismer, Wood, & Cota, 2014, H. dushanensis (Zhou & Liu, 1981), H. flaviventris, H. huishuiensis Yan, Lin, Guo, Li, & Zhou, 2016, H. jinpingensis (Zhou & Liu, 1981), H. khlonglanensis, H. kiziriani, H. linnwayensis, H. longlingensis (Zhou & Liu, 1981), H. montawaensis Grismer, Wood, Thura, Zin, Quah, Murdoch, Grismer, Li, Kyaw, & Lwin, 2017, H. tonywhitteni Grismer, Wood, Thura, Zin, Quah, Murdoch, Grismer, Li, Kyaw, & Lwin, 2017, H. yunnanensis, and H. zugi Nguyen, Lehmann, Le, Duong, Bonkowski, Ziegler, 2013]; infralabial scales 12 (versus ≤10 in H. changningensis, H. linnwayensis, H. montawaensis, and H. tonywhitteni); dorsal scales within one orbital diameter 30 (versus ≤27 in H. aurantiacus, H. banaensis, H. changningensis, H. chiangmaiensis, H. flaviventris, H. huishuiensis, H. khlonglanensis, H. kiziriani, H. linnwayensis, H. montawaensis, H. tonywhitteni, H. yunnanensis, and H. zugi); ventral scales within one orbital diameter 11 (versus ≤9 in H. changningensis, H. huishuiensis, H. linnwayensis, H. montawaensis, H. tonywhitteni, and 14–16 in H. zugi); total lamellae on hand 18 (versus ≤16 in H. aurantiacus, H. changningensis, H. chiangmaiensis, H. dushanensis, H. flaviventris, H. jinpingensis, H. khlonglanensis, H. kiziriani, H. linnwayensis, H. longlingensis, H. montawaensis, H. yunnanensis, and H. zugi); total lamellae on foot 19 (versus ≤16 in H. aurantiacus, H. changningensis, H. chiangmaiensis, H. khlonglanensis, H. longlingensis, and H. yunnanensis); subdigital lamellae on Finger I four (versus three in H. huishuiensis, H. tonywhitteni and ≥5 in H. banaensis, H. dushanensis, H. flaviventris, H. jinpingensis, H. khlonglanensis, H. kiziriani, and H. longlingensis); subdigital lamellae on Toe I five (versus ≤4 in H. changningensis, H. chiangmaiensis, H. huishuiensis, H. montawaensis, H. tonywhitteni, and seven in H. dushanensis); and continuous precloacofemoral pores 18 (versus ≥20 in H. dushanensis, H. jinpingensis, H. tonywhitteni, and ≤13 in H. flaviventris, H. kiziriani). 
CommentHabitat: the holotype was collected approximately 1 m above the ground inside of a vertical palm tree in wet evergreen forest during the morning (Fig. 1B).

Only known from the holotype. 
EtymologyThe specific epithet is taken from the Latin indus for belonging to India, and sobrinus for maternal cousin, in reference to its close relationship, along with H. flaviventris, in maternally-inherited mitochondrial sequence data to the Indian endemic H. aurantiacus (Fig. 2; Grismer et al. 2013, 2014a,b, 2015, 2017; Nguyen et al. 2013, 2014; Ngo et al. 2014; Yan et al. 2016; Sukprasert et al. 2018). 
References
  • ELIADES, SAMUEL J.; SOMPHOUTHONE PHIMMACHAK, NIANE SIVONGXAY, CAMERON D. SILER, BRYAN L. STUART 2019. Two new species of Hemiphyllodactylus (Reptilia: Gekkonidae) from Laos. Zootaxa 4577 (1): 131–147 - get paper here
  • NGUYEN, TRUONG QUANG; QUYEN HANH DO, HANH THI NGO, ANH VAN PHAM,<br />CUONG THE PHAM, MINH DUC LE, & THOMAS ZIEGLER 2020. Two new species of Hemiphyllodactylus (Squamata: Gekkonidae) from Hoa Binh Province, Vietnam. Zootaxa 4801 (3): 513–536 - get paper here
 
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