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Hemiphyllodactylus khlonglanensis SUKPRASERT, SUTTHIWISES, LAUHACHINDA & TAKSINTUM, 2018

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Common NamesE: Khlong Lan Dwarf Gecko
Thai: Ching Chok Khao Sung Khlong Lan 
SynonymHemiphyllodactylus khlonglanensis SUKPRASERT, SUTTHIWISES, LAUHACHINDA & TAKSINTUM 2018 
DistributionW Thailand (Kamphaeng Phet)

Type locality: Khlong Lan District, Kamphaeng Phet Province, western Thailand  
Reproductionoviparous (not imputed, fide Zimin et al. 2022) 
TypesHolotype: ZMKU TM001043N (Kasetsart University, Thailand), adult male, collected by Apisak Sukprasert on May 8, 2016. Paratypes. Seven specimens from the same locality as the holotype. ZMKU TM000999N–ZMKU TM001004N collected on December 25, 2015 and ZMKU TM001044N collected on May 8, 2016. 
DiagnosisHemiphyllodactylus khlonglanensis sp. nov. may be diagnosed by the combination of characteristics: (1) a bisexual taxon; (2) a maximum SVL of 43.05 mm in males and 43.14 mm in females; (3) 7–10 chin scales; (4) 5 circumnasal scales; (5) 2–4 scales between supranasals; (6) 9–12 supralabials; (7) 8–11 infralabials; (8) 10–15 dorsal scale rows; (9) 7–10 ventral scale rows; (10) lamellar formula on forefoot 3-4-4-4 or 4-4-4-4; (11) lamellar formula on hindfoot 4-4-4-4; (12) precloacal and femoral pores in a continuous series, 19–20 pores in males; (13) 1 cloacal spur on each side present in both sexes; (14) dark postorbital stripe; and (15) anteriorly projecting arms of postsacral mark.

Comparisons. Hemiphyllodactylus khlonglanensis sp. nov. is placed within a clade (lineage A; Figure 2) along with four of its congeners. It has a lamellar formula of 3-4-4-4 or 4-4-4-4 on the forefoot and 4-4- 4-4 on the hindfoot, which separates it from H. changningensis Guo, Zhou, Yan & Li (3-3/4-3/4-3 and 3-4-4-4 or 3- 3-3-3, respectively) and H. chiangmaiensis (3-3-3-3 or 3-4-3-3 and 3-3-3-3 or 3-4-4-4, respectively). Hemiphyllodactylus khlonglanensis sp. nov. has five or six lamellae on its first finger in contrast with H. changningensis and H. chiangmaiensis which have three or four and H. jinpingensis which has four, and it has five subdigital lamellae on the first toe, whereas H. changningensis and H. chiangmaiensis have three or four. The postsacral mark in H. khlonglanensis sp. nov. has anteriorly projecting arms, which are lacking in H. longlingensis Zhou & Liu and H. jinpingensis. The maximum known SVL of H. khlonglanensis sp. nov. is 43.1 mm, whereas in H. jinpingensis is 39.6 mm. Diagnostic characteristics of Hemiphyllodactylus are shown in Table 4 in SUKPRASERT et al. 2018. 
CommentDistribution: for a map see Grismer et al. 2020: 172 (Fig. 1). 
EtymologyThis specific epithet is derived from the name of Khlong Lan District which refers to the type locality. 
  • GRISMER, L. L., YUSHCHENKO, P. V., PAWANGKHANANT, P., NAIDUANGCHAN, M., NAZAROV, R. A., ORLOVA, V. F., ... & POYARKOV, N. A. 2020. A new species of Hemiphyllodactylus Bleeker (Squamata; Gekkonidae) from Peninsular Thailand that converges in morphology and color pattern on Pseudogekko smaragdinus (Taylor) from the Philippines. Zootaxa 4816 (2): 171-190 - get paper here
  • NGUYEN, TRUONG QUANG; QUYEN HANH DO, HANH THI NGO, ANH VAN PHAM,<br />CUONG THE PHAM, MINH DUC LE, & THOMAS ZIEGLER 2020. Two new species of Hemiphyllodactylus (Squamata: Gekkonidae) from Hoa Binh Province, Vietnam. Zootaxa 4801 (3): 513–536 - get paper here
  • SUKPRASERT, APISAK; SURAPOP SUTTHIWISES, VIRAYUTH LAUHACHINDA, WUT TAKSINTUM 2018. Two new species of Hemiphyllodactylus Bleeker (Squamata: Gekkonidae) from Thailand. Zootaxa 4369 (3): 363–376 - get paper here
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