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Hemiphyllodactylus titiwangsaensis ZUG, 2010

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Common NamesE: Titiwangsa slender gecko 
SynonymHemiphyllodactylus titiwangsaensis ZUG 2010: 48
Hemiphyllodactylus titiwangsaensis — GRISMER 2011
Hemiphyllodactylus titwangsaensis — GRISMER & QUAH 2019 (in error) 
DistributionW Malaysia (Pahang Province)

Type locality: Malaysia, Pahang Province, Cameron Highlands, Gunong Brinchang (= Berincang) summit area.  
Reproductionoviparous (not imputed, fide Zimin et al. 2022) 
TypesHolotype: ZRC 2.4782, adult male, collected by H. H. Tan and others, 25 June 2000. Paratypes: ZFMK, ZRC 
CommentMajor diagnostic features: bisexual taxon; caecum and gonadal ducts not pigmented; pre- cloacal–femoral pore series continuous in males (TotPore 17–39), absent in females; chin scales bordering mental and first infralabial distinctly enlarged; digital lamellae formulae usually 3-4-4-4 (forefoot) and 4-4-5-5 or 4-5-5-5 (hindfoot); average adult SVL ~49 mm; dorsal and lateral trunk pattern of dark brown irregular transverse bands, muted postsacral bar of narrow white arms onto hips.

Behavior: nocturnal

Distribution: see map in Grismer et al. 2015 (Fig. 1C). 
EtymologyThese geckos occur in the south central region of the Banjaran Titiwangsa; hence the taxon is a resident (likely endemic) of Titiwangsa and so named. 
  • Böhme, Wolfgang 2014. Herpetology in Bonn. Mertensiella 21. vi + 256 pp. - get paper here
  • COBOS, ANTHONY; L. LEE GRISMER, PERRY L. WOOD, JR., EVAN S. H. QUAH, SHAHRUL ANUAR, MOHD ABDUL MUIN 2016. Phylogenetic relationships of geckos of the Hemiphyllodactylus harterti group, a new species from Penang Island, Peninsular Malaysia, and a likely case of true cryptic speciation. Zootaxa 4107 (3): 367-380 - get paper here
  • GOLDBERG, S. R., C. R. BURSEY, AND L. L. GRISMER 2020. Hemiphyllodactylus titiwangsaensis (Titiwangsa Slender Gecko). Endoparasites. Herpetological Review 51: 600.
  • Goldberg, Stephen R. and L. Lee Grismer. 2014. Hemiphyllodactylus titiwangsaensis (Titiwangsa slender gecko) reproduction. Herpetological Review 45 (3): 500 - get paper here
  • GRISMER, L. LEE; & EVAN S. H. QUAH 2019. An updated and annotated checklist of the lizards of Peninsular Malaysia, Singapore, and their adjacent archipelagos. Zootaxa 4545 (2): 230–248 - get paper here
  • GRISMER, L. LEE; AWAL RIYANTO, DJOKO T. ISKANDAR & JIMMY A. MCGUIRE 2014. A new species of Hemiphyllodactylus Bleeker, 1860 (Squamata: Gekkonidae) from Pulau Enggano, southwestern Sumatra, Indonesia. Zootaxa 3821 (4): 485–495 - get paper here
  • Grismer, L. Lee; Perry L. Wood Jr, Shahrul Anuar, Evan S. H. Quah, Mohd Abdul Muin, Chan Kin Onn, Alexandra X. Sumarli and Ariel I. Loredo 2015. Repeated evolution of sympatric, palaeoendemic species in closely related, co-distributed lineages of Hemiphyllodactylus Bleeker, 1860 (Squamata: Gekkonidae) across a sky-island archipelago in Peninsular Malaysia. Zoological Journal of the Linnean Society 174 (4): 859–876; DOI: 10.1111/zoj.12254 - get paper here
  • Grismer, L.L. 2011. Lizards of Peninsular Malaysia, Singapore and their adjacent archipelagos. Edition Chimaira, Frankfurt, 728 pp. [review in Herp. Rev. 43: 155] - get paper here
  • Grismer, L.L.; PERRY L. WOOD Jr, SHAHRUL ANUAR, MOHD ABDUL MUIN, EVAN S. H. QUAH, JIMMY A. MCGUIRE,, RAFE M. BROWN,0, NGO VAN TRI and PHAM HONG THAI 2013. Integrative taxonomy uncovers high levels of cryptic species diversity in Hemiphyllodactylus Bleeker, 1860 (Squamata: Gekkonidae) and the description of a new species from Peninsular Malaysia. Zoological Journal of the Linnean Society 169: 849–880 - get paper here
  • NGO VAN TRI; L. LEE GRISMER, PHAM HONG THAI & P. L. WOOD, JR. 2014. A new species of Hemiphyllodactylus Bleeker, 1860 (Squamata: Gekkonidae) from Ba Na–Nui Chua Nature Reserve, Central Vietnam. Zootaxa 3760 (4): 539–552 - get paper here
  • NGUYEN, TRUONG QUANG; QUYEN HANH DO, HANH THI NGO, ANH VAN PHAM,<br />CUONG THE PHAM, MINH DUC LE, & THOMAS ZIEGLER 2020. Two new species of Hemiphyllodactylus (Squamata: Gekkonidae) from Hoa Binh Province, Vietnam. Zootaxa 4801 (3): 513–536 - get paper here
  • Zimin, A., Zimin, S. V., Shine, R., Avila, L., Bauer, A., Böhm, M., Brown, R., Barki, G., de Oliveira Caetano, G. H., Castro Herrera, F., Chapple, D. G., Chirio, L., Colli, G. R., Doan, T. M., Glaw, F., Grismer, L. L., Itescu, Y., Kraus, F., LeBreton 2022. A global analysis of viviparity in squamates highlights its prevalence in cold climates. Global Ecology and Biogeography, 00, 1–16 - get paper here
  • Zug, G. R. 2010. Speciation and Dispersal in a Low Diversity Taxon: The Slender Geckos Hemiphyllodactylus (Reptilia, Gekkonidae). Smithsonian contributions to zoology (631): 1-70 - get paper here
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