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Heteronotia fasciolata PEPPER, DOUGHTY, FUJITA, MORITZ & KEOGH, 2013

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Common NamesE: Pale-headed Gecko 
SynonymHeteronotia fasciolatus PEPPER, DOUGHTY, FUJITA, MORITZ & KEOGH 2013
Heteronotia fasciolata — COGGER 2014: 976 
DistributionAustralia (NT)

Type locality: Harts Range racecourse, central Australia, NT (-23.02284°S, 134.93135°E),  
TypesHolotype: NTM R36284 (male), collected on 3 June 2010 by M. Fujita, M. Pepper, and C. Moritz. Paratypes. — MCZ 188177 Harts Range racecourse (-23.02284°S, 134.93135°E); MCZ 188183 (male) Tylers Pass, NT (-23.62938°S, 132.35660°E); MCZ 188182 (male), Ormiston Gorge, NT (-23.63799uS, 132.73291uE); NTM R36288 (female)
Tylers Pass, NT (-23.62938°S, 132.35660°E); NTM R36297 (male), Ormiston Gorge, NT (223.63799°S, 132.73291°E). 
DiagnosisDiagnosis. — Distinguished from congeners by medium body size (to 57.0 mm SVL), slightly robust habitus, moderately wide head (to 13.7 mm HeadW), tail stout at base, small tympanum, typically 8 or 9 supralabials and 6 or 7 infralabials, enlarged dorsal tubercles surrounded by at least one smaller granule anterior and posterior to scale, and usually two smaller granules to sides, dorsum with 6–8 strongly contrasting bands; edges of bands with dark brown border and irregular edge (some bands breaking up), top of head pale.

Comparison to other species. — Heteronotia fasciolatus sp. nov. can be distinguished from H. spelea and H. atra sp. nov. by the strongly contrasting dorsal bands being uneven and often broken, and never straight edged. Furthermore, the head is broad and pale, with light red or brown pigment. The enlarged dorsal tubercles are widely spaced compared to H. spelea and H. atra sp. nov., generally with two small granular scales separating them. It can be further differentiated from H. atra sp. nov. by having a small ear opening. 
CommentHabitat. — Sheltering among rocks naturally, but can be found among human-made structures such as sheet metal in rocky surrounds.

Similar species: H. binoei (see sympatry)

Sympatry: the CA6 chromosome race of H. binoei in central Australia. Here H. binoei also posess a banded morphology, with occasional samples strikingly similar to H. fasciolatus. 
EtymologyIn a PhD thesis, Stewart (1996) informally proposed the name H. fasciolatus based on four karyotyped specimens from Undoolya Station in Central Australia. Fasciolatus is a diminutive of the Latin fascia, meaning ‘band’. 
  • Cogger, H. G. 2014. Reptiles and Amphibians of Australia, 7th ed. CSIRO Publishing, xxx + 1033 pp. - get paper here
  • Pepper, Mitzy; Paul Doughty, Matthew K. Fujita, Craig Moritz, J. Scott Keogh 2013. Speciation on the Rocks: Integrated Systematics of the Heteronotia spelea Species Complex (Gekkota; Reptilia) from Western and Central Australia. PLoS One 8 (11): e78110. doi:10.1371/journal.pone.0078110 - get paper here
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