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Hypnale zara (GRAY, 1849)

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Higher TaxaViperidae, Crotalinae, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies 
Common Names 
SynonymTrigonocephalus zara GRAY 1849
Hypnale zara — MADUWAGE et al. 2009
Hypnale zara — WALLACH et al. 2014: 343 
DistributionSri Lanka

Type locality: see comment  
Reproductionovoviviparous 
TypesHolotype: BMNH 1946. 1. 19. 96 male, 303 mm SVL, coll. “Mr Argent” 
DiagnosisDiagnosis: Hypnale zara is distinguished from its congeners by a combination of the following characters: tip of snout distinctly elevated; 10–19 minute scales form a wart-like protuberance on snout tip (Fig. 6a, b); 18–39 heterogeneous, small scales on internasal-prefrontal region (Fig. 6b); scales around eye 6 (supraocular + 2 postoculars + 2 preoculars + postfoveal) (Fig. 6a); maxillary pit surrounded by 3 scales (lower preocular + postfoveal + second supralabial) (Fig. 6a); 2 postoculars, the upper one small, the lower one reaching postfoveal; no scales between postfoveal and 3rd supralabial (Fig. 6a); lower postocular, 1st lower temporal, 4th and 5th supralabials in contact, no scale between them. Supralabials 7 or 8; lacunal scale absent; 2nd supralabial extends upward to reach loreal; costal scales keeled (Fig. 6g); ventrals 134–157; 34– 51 scales in subcaudal series, divided; hemipenial lobes not spinous (Fig. 6d). Hypnale zara is distinguished from H. hypnale by having the tip of the snout distinctly elevated (vs snout tip not raised); possessing (vs lacking) a wart-like protuberance on the snout tip; having 18–39 (vs 5–15) scales on internasal-prefrontal region; the maxillary pit surrounded by 3 (vs 4–5) scales; and the lacunal scale absent (vs present). It is distinguished from H. nepa by having the tip of the snout distinctly elevated (vs flattened or only slightly raised); 18–39 (vs 12–25) scales on the internasal-prefrontal region; the maxillary pit surrounded by 3 (vs 4–5) scales; no (vs 1–3) scales between postfoveal and 3rd supralabial; lower postocular, 1st lower temporal, 4th and 5th supralabials in contact (vs separated by a rhomboid scale of similar size); lacunal scale absent (vs present); all costal scales keeled (vs mid-dorsal scales smooth); 134–157 (vs122–134) ventrals; and having the hemipenial lobes not spinous (vs with large spines on the proximal two-thirds of their length). Hypnale zara is distinguished from Hypnale sp. ‘amal’ by having the tip of the snout distinctly elevated (vs not elevated); possessing (vs lacking) a wart-like protuberance on the snout tip; having 18–39 (vs 10) scales on internasal-prefrontal region; the maxillary pit surrounded by 3 (vs 4) scales; 7 or 8 (vs 9) supralabials; the lacunal scale absent (vs present); first infralabials on either side separated from each other by a median suture (vs medially fused); and two rows of distinct, bilaterally symmetrical blotches meeting on dorsal midline (vs a distinct purple-brown paravertebral stripe about 3 scales wide extending from neck to level of vent). (Maduwage et al. 2009: 15)


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CommentVenomous!

The original Type locality was given as “Singapore” which is obviously wrong. 
References
  • Ganesh, S. R.; Subramanian Bhupathy, Patrick David, N. Sathishkumar, G. Srinivas 2014. Snake Fauna of High Wavy Mountains, Western Ghats, India: Species Richness, Status, and Distribution Pattern. Russ. J. Herpetol. 21 (1): 53-64
  • Gray, J. E. 1849. Catalogue of the specimens of snakes in the collection of the British Museum. Edward Newman, London, i-xv; 1-125. - get paper here
  • Janzen, Peter and Malaka Bopage 2011. The herpetofauna of a small and unprotected patch of tropical rainforest in Morningside, Sri Lanka. Amphibian & Reptile Conservation 5 (2): 1-13 - get paper here
  • Karunarathna, Suranjan D. M. S. and A. A. Thasun Amarasinghe 2011. A PRELIMINARY SURVEY OF THE REPTILE FAUNA IN NILGALA FOREST AND ITS VICINITY, MONARAGALA DISTRICT, SRI LANKA. Taprobanica 3 (2): 69-76 - get paper here
  • Madawala, Majintha; Thilina D Surasinghe, Anslem De Silva, Dinesh Gabadage, Madhava Botejue, Indika Peabotuwage, Dushantha Kandambi, Suranjan Karunarathna 2019. Reappraisal of Herpetofauna Recorded from Jaffna Peninsula in Northern Sri Lanka with Remarks on Conservation, Diversity, and Distribution. Russ. J. Herpetol. 26 (5): 247-260 - get paper here
  • MADUWAGE, KALANA; ANJANA SILVA, KELUM MANAMENDRA-ARACHCHI & ROHAN PETHIYAGODA 2009. A taxonomic revision of the South Asian hump-nosed pit vipers (Squamata: Viperidae: Hypnale). Zootaxa 2232: 1-28 - get paper here
  • Pyron, R. Alexander; H.K. Dushantha Kandambi, Catriona R. Hendry, Vishan Pushpamal, Frank T. Burbrink, Ruchira Somaweera 2013. Genus-level phylogeny of snakes reveals the origins of species richness in Sri Lanka. Molecular Phylogenetics and Evolution 66 (3): 969-978 - get paper here
  • Samarawickrama, V.A.M.P.K.; H.I.G.C. Kumara, D.R.N.S. Samarawickrama 2019. Diversity of Reptiles in the Eastern and Southern parts of the Sinharaja Rain Forest. Journal of Tropical Forestry and Environment - get paper here
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp.
 
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