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Lampropeltis triangulum (LACÉPÈDE, 1789)

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Higher TaxaColubridae, Colubrinae, Lampropeltini, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
SubspeciesLampropeltis triangulum gentilis (BAIRD & GIRARD 1853)
Lampropeltis triangulum syspila (COPE 1889)
Lampropeltis triangulum triangulum (LACÉPÈDE 1789) 
Common NamesE: Eastern Milksnake
E: Louisiana Milksnake [amaura]
E: Red Milksnake [syspila]
G: Rote Königsnatter, Milchschlange, (Östliche) Dreiecksnatter
E: Mexican Milksnake [annulata]
E: Central Plains Milksnake [gentilis]
E: Pale Milksnake [multistriata]
E: Utah Milksnake [taylori]
E: New Mexico Milksnake [celaenops]
S: Barila 
SynonymColuber triangulum LACÉPÈDE 1789: 86
Coluber doliatus — SHAW & NODDER 1795: plate 254
Coluber eximius HARLAN 1827
Ophibolus clericus BAIRD & GIRARD 1853
Ablabes triangulum — DUMÉRIL, BIBRON & DUMÉRIL 1854: 315 (?)
Lampropeltis triangula — COPE 1860: 256
Lampropeltis amaura COPE 1861: 258 (footnote)
Coronella eximia — JAN 1866 ?
Ophibolus doliatus amaura — COPE 1875: 36 (part.)
Ophibolus triangulus — GARMAN 1884: 66
Ophibolus doliatus parallelus COPE 1889: 385
Ophibolus doliatus collaris COPE 1889: 383
Coronella annulata — GÜNTHER 1893 (fide VILLA et al. 1988)
Ophibolus doliatus annulatus — COPE 1893: 1067
Ophibolus doliatus parallelus — COPE 1893: 1067
Ophibolus doliatus collaris — COPE 1893: 1068
Ophibolus doliatus clericus — COPE 1893: 1068
Coronella triangulum collaris — BOETTGER 1898: 72
Osceola doliata parallela — COPE 1900: 893
Osceola doliata triangula — COPE 1900: 885
Osceola doliata collaris — COPE 1900: 886
Osceola doliata clerica — COPE 1900: 888
Osceola doliata parallela — COPE 1900: 893
Osceola doliata triangula — MCATEE 1907
Lampropeltis triangulum amaura — STEJNEGER & BARBOUR 1917: 90 (part.)
Lampropeltis polyzonus — DITMARS 1934 (fide VILLA et al. 1988)
Lampropeltis triangulum amaura — BURT 1935 (part.)
Lampropeltis polyzona blanchardi — SCHMIDT & ANDREWS 1936 (VILLA et al.)
Lampropeltis doliata amaura — KLAUBER 1948: 11 (part.)
Lampropeltis triangulum — STEBBINS 1985: 194
Lampropeltis triangulum triangulum — CONANT & COLLINS 1991: 207
Lampropeltis triangulum amaura — CONANT & COLLINS 1991: 210 (part.)
Lampropeltis triangulum — LINER 1994
Lampropeltis triangulum triangulum — CROTHER 2000: 64
Lampropeltis triangulum amaura — CROTHER 2000: 64 (part.)
Lampropeltis triangulum amaura — TENNANT & BARTLETT 2000: 434 (part.)
Lampropeltis triangulum triangulum — TENNANT & BARTLETT 2000: 425
Lampropeltis triangulum triangulum — CROTHER et al. 2012
Lampropeltis triangulum — CROTHER et al. 2012
Lampropeltis triangulum amaura — CROTHER et al. 2012 (part.)
Lampropeltis triangulum — RUANE et al. 2014
Lampropeltis triangulum — WALLACH et al. 2014: 359
Lampropeltis triangulum triangulum — GUYER et al. 2018
Lampropeltis triangulum — CHAMBERS et al. 2022

Lampropeltis triangulum gentilis (BAIRD & GIRARD 1853)
Ophibolus gentilis BAIRD & GIRARD 1853: 90
Lampropeltis annulata COPE 1860: 257 (refers to ms. of KENNICOTT) (part.)
Lampropeltis annulata KENNICOTT 1861: 329 (part.)
Lampropeltis multistriata KENNICOTT 1861: 328
Lampropeltis multistrata — COPE 1861: 566
Coronella doliata var. gentilis — JAN 1866
Ophibolus multistratus — COPE 1875: 36
Ophibolus triangulus var. gentilis — GARMAN 1884: 66
Ophibolus multistratus — GARMAN 1887: 128
Lampropeltis multistrata – STEJNEGER 1891: 502
Ophibolus doliatus gentilis — COPE 1893: 1068
Osceola doliata gentilis — COPE 1900: 894
Ophibolus multistratus — COPE 1900: 909
Osceola doliata coccinea — COPE 1900: 896 (?)
Lampropeltis pyrrhomelana celaenops STEJNEGER 1902: 153
Lampropeltis pyromelana celaenops STEJNEGER & BARBOUR 1917: 89
Coronella multistrata – WERNER 1929: 129
Lampropeltis doliata multistrata — TANNER & LOOMIS 1957: 12
Lampropeltis doliata celaenops — TANNER & LOOMIS 1957: 15
Lampropeltis doliata taylori TANNER & LOOMIS 1957: 31
Lampropeltis triangulum celaenops — STEBBINS 1966: 16
Lampropeltis triangulum multistrata — STEBBINS 1966: 161
Lampropeltis triangulum gentilis — MEHRTENS 1987: 127
Lampropeltis triangulum multistriata — CONANT & COLLINS 1991: 209
Lampropeltis triangulum celaenops — CONANT & COLLINS 1991: 209
Lampropeltis triangulum gentilis — CONANT & COLLINS 1991: 209
Lampropeltis triangulum annulata — CONANT & COLLINS 1991: 209 (part.)
Lampropeltis taylori — COLLINS 1991: 43
Lampropeltis triangulum taylori — WILLIAMS 1994
Lampropeltis triangulum multistrata — WILLIAMS 1994
Lampropeltis triangulum celaenops — WILLIAMS 1994
Lampropeltis triangulum multistriata — CROTHER 2000: 64
Lampropeltis triangulum gentilis — CROTHER 2000: 64
Lampropeltis triangulum celaenops — CROTHER 2000: 64
Lampropeltis triangulum taylori — CROTHER 2000: 64
Lampropeltis triangulum annulata — CROTHER 2000: 64
Lampropeltis triangulum multistrata [sic] — TENNANT & BARTLETT 2000: 442
Lampropeltis triangulum annulata — TENNANT & BARTLETT 2000: 436 (part.)
Lampropeltis triangulum celaenops — TENNANT & BARTLETT 2000: 438
Lampropeltis triangulum gentilis — TENNANT & BARTLETT 2000: 440
Lampropeltis triangulum gentilis — BARTS 2006
Lampropeltis triangulum multistriata — CROTHER et al. 2012
Lampropeltis triangulum celaenops — CROTHER et al. 2012
Lampropeltis triangulum gentilis — CROTHER et al. 2012
Lampropeltis triangulum taylori — CROTHER et al. 2012
Lampropeltis triangulum annulata — CROTHER et al. 2012 (part.)
Lampropeltis gentilis — RUANE et al. 2014
Lampropeltis triangulum gentilis — CHAMBERS et al. 2022

Lampropeltis triangulum syspila (COPE 1889)
Ophibolus doliatus syspilus COPE 1889: 381 (part.)
Lampropeltis doliatus syspilus — HAY 1892: 516 (part.)
Ophibolus doliatus syspilus — COPE 1893: 1067
Osceola doliata syspila — COPE 1900: 891 (part.)
Lampropeltis triangulum syspila — RUTHVEN 1919: 3 (part.)
Lampropeltis triangulum syspila — BURT 1935 (part.)
Lampropeltis triangulum syspila — MEHRTENS 1987: 128
Lampropeltis triangulum syspila — CONANT & COLLINS 1991: 208 (part.)
Lampropeltis triangulum syspila — CROTHER 2000: 64 (part.)
Lampropeltis triangulum syspila — TENNANT & BARTLETT 2000: 432 (part.)
Lampropeltis triangulum syspila — CROTHER et al. 2012 (part.)
Lampropeltis triangulum syspila — GUYER et al. 2018
Lampropeltis triangulum syspila — CHAMBERS et al. 2022 
DistributionCanada (Ontario, S Quebec),
USA (N Texas, W Oklahoma, C/W Kansas, E Colorado, and SC / SW Nebraska, Illinois, Minnesota, Michigan, Iowa, Missouri, Tennessee, Arkansas, N Louisiana, N Mississippi, N Alabama, N Georgia, N South Carolina, N North Carolina, Virginia, West Virginia, Kentucky, Indiana, Illinois, Wisconsin, Ohio, Maryland, Pennsylvania, New Jersey, New York, Rhode Island, Connecticut, Massachusetts, New Hampshire, Vermont, S Maine)
Mexico (Coahuila)

Type locality: “America”

amaura (invalid): SW Arkansas, SE Oklahoma, E Texas and Louisiana. Type locality, original entry of type reads "?Mississippi," restricted by Schmidt (1953) to the vicinity of New Orleans, Louisiana. Williams (1978) suggested that the type locality be considered as unknown, inasmuch as the characteristics of the holotype do not agree with data taken from specimens from New Orleans and vicinity.

celaenops (invalid): USA (SW Texas, E New Mexico, SE Arizona). Type locality: Mesilla Valley, New Mexico.

gentilis: Kansas, W Oklahoma, E Colorado, Texas panhandle, C/SW Nebraska, C/W Kansas; Type locality: "north fork of the Red River, near Sweetwater Creek, Wheeler County, Texas."

multistrata (invalid): USA (C/NW Nebraska, S Montana, SW North Dakota, N/C Wyoming); Type locality: "Fort Lookout, Nebraska," now in Lyman County, South Dakota.

syspila: USA (Nebraska, Oklahoma, Kansas)

taylori (invalid): C/NE Utah, W Colorado, N Arizona; Type locality: "from approximately 2 miles north of Alpine, Utah County, Utah."  
Reproductionoviparous. KREUTZ (2005) reports hybrids between E. guttata and Lampropeltis getulus californiae, between E. guttata and L. pyromelana, and between E. guttata and L. triangulum sinaloae/nelsoni! 
TypesHolotype: lost or never designated (fide WILLIAMS 1994)
Holotype: USNM 5282, an adult male [amaura]
Syntypes: USNM 2192 etc [Ophibolus doliatus collaris]
Holotype: USNM 13380 [Ophibolus doliatus syspilus]
Holotype: USNM 10544 [Ophibolus doliatus parallelus]
Holotype: USNM 2380 [Ophibolus clericus]
Lectotype: USNM 131737 (old catalog number 1852 or 1853), adult male (designated by BLANCHARD 1921) [gentilis]
Holotype: USNM 22375 [celaenops]
Holotype: USNM 1842, young male [multistriata]
Holotype: BYU 10533, adult female [taylori] 
DiagnosisDiagnosis: Lampropeltis triangulum differs from L. rutheni, L. mexicana, and L. alterna in having the head less distinct from the body. In addition, this species differs from L. ruthveni (within the range of the latter) in having less uniform red rings, often obscured by black pigment middorsally. Another difference between L. triangulum on the one hand, and ruthveni and L. mexicana on the other, is that in the latter two the black rings are edged with a paler color (usually a pale lime green). Lampropeltis ruthveni has a lower ventral count (182-196) than sympatric L. triangulum (196-221). All members of the mexicana group tend to have considerable gray in the body colors, such is not so in triangulum, except some triangulum in the northeastern United States. Lampropeltis triangulum differs from L. zonata in that the white rings widen laterally in the former; from L. pyromelana in having a black, black and white, or red snout, rather than white. Lampropeltis pryomelana has 46-85 white rings on body, more than L. triangulum (10-32). Both L. pyromelana and L. zonata have a higher ventral scute number (213-238, and 194-227, respectively) than any of the adjacent western United States members of L. triangulum (174-208). Lampropeltis triangulum differs from both L. calligaster and L. getula in having an alternating pattern of white (yellow), red and black rings or blotches, the latter reaching the ventrals or nearly so, and in having the posterior two maxillary teeth longer and stouter than the preceding teeth. (Williams 1994)


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CommentSynonymy after PETERS & OREJAS-MIRANDA 170, WILLIAMS 14, and RUANE et al. 2014, and Chalmers et al. 2022 (although they did not comment on synonyms beyond moving gentilis back to triangulum as subspecies).

Distribution: L. gentilis is not in Nuevo León but annulata is, fide Nevárez-de los Reyes et al. 2016. See map in Chalmers et al. 2022: 359 (Fig. 1).

Distribution: revised after genetic analysis of Ruane et al. 2014. Previously, this species was also reported from Mexico (e.g. Jalisco, San Luis Potosí, Nuevo Léon, Coahuila, Nayarit, Sonora etc., e.g. by Lemos-Espinal et al. 2016), south to Colombia (Valle del Cauca, Valencia et al. 2014).

Subspecies: Lampropeltis triangulum elapsoides (HOLBROOK 1838) has been elevated to species status. Ruane et al. (2014) elevated a number of subspecies of L. triangulum to full species and synonymized the remaining subspecies as follows: Subspecies synonymized under L. micropholis include L. t. gaigeae and L. t. andesiana. Subspecies synonymized under L. abnorma include L. t. blanchardi, L. t. hondurensis, L. t. oligozona and L. t. stuarti. Subspecies synonymized under L. polyzona include L. t. arcifera, L. t. conanti, L. t. campbelli, L. t. nelsoni, L. t. smithi, and L. t. sinaloae. Subspecies synonymized under L. annulata include L. t. dixoni. Subspecies synonymized under L. gentilis include L. t. celaenops, L. t. multistriata, L. t. taylori, L. t. amaura (in part), L. t. syspila (in part) and L. t. annulata (in part). Subspecies synonymized under L. triangulum include L. t. syspila (in part) and L. t. amaura (in part). L. gentilis and triangulum have a broad contact zone with little reproductive isolation which does not warrant recognition as full species fide Chalmers et al. 2022. Basett & Pandelis (2024) did not accept the separate species model of Ruane et al. 2014 either.

Nomenclature: LACÉPÈDE’s name conserved through ICZN opinion 1463. Coluber doliatus was suppressed via Opinion 804. The original spelling of multistriata was apparently a misspelling of multistrata, fide Cope 1861: 566, Williams 1994: 7.

Mimicry: Lampropeltis triangulum mimicks Micrurus bernadi, M. limbatus and M. diastema (TORRE-LORANCA et al. 2006).

NCBI taxonID: 165043 [gentilis] 
EtymologyThe name triangulum is derived from the Latin meaning "triangle," in reference to the shape of the anterior edge of the first blotch.

The name gentilis is derived from the Latin, meaning "of the same race or clan", presumably in reference to its relationship to other members of the species.

The name celaenops may be derived from the Greek word celaeno, which means "black or dark," and the ending -ops, meaning eye.

The name multistrata is derived from the Latin multus, meaning "many," and the Latin stratum, meaning "layer," apparently in reference to the number of dorsal scale rows present.

The name taylori isa patronym honoring Edward H. Taylor, designated "our friend and teacher" by describers. 
References
  • Aguilar-López, José Luis and Eduardo Pineda 2013. A contribution to our knowledge of the false coral snake’s (Lampropeltis triangulum, Lacépède 1788) diet. Herpetology Notes 6: 89-90
  • Akcali, C. K. and Pfennig, D. W. 2017. Geographic variation in mimetic precision among different species of coral snake mimics. J. Evol. Biol., doi:10.1111/jeb.13094 - get paper here
  • Allen, Morrow J. 1932. A survey of the Amphibians and reptiles of Harrison County, Mississippi. American Museum Novitates (542): 1-20 - get paper here
  • Anonymous 2007. Snakes of New Jersey. New Jersey Division of Fish and Wildlife
  • Applegate, Robert W. 1988. The Pueblan milksnake (Lampropeltis triangulum campbelli): a case for captive propagation. Vivarium 1 (2): 49-53
  • Arenas-Monroy, J.C., R. A. Carbajal-Márquez. 2012. Lampropeltis triangulum. MÉXICO: GUANAJUATO. Herpetological Review 43: 106 - get paper here
  • Armstrong, M.P., D. Frymire & E.J. Zimmerer 2001. Analysis of Sympatric Populations of Lampropeltis triangulum syspila and Lampropeltis triangulum elapsoides, in Western Kentucky and Adjacent Tennessee with Relation to the Taxonomic Status of the Scarlet Kingsnake Journal of Herpetology 35 (4): 688-693. - get paper here
  • Augstenová, Barbora; Sofia Mazzoleni, Lukáš Kratochvíl Orcid and Michail Rovatsos 2017. Evolutionary Dynamics of the W Chromosome in Caenophidian Snakes. Genes 2018, 9(1), 5; doi:10.3390/genes9010005 - get paper here
  • Badman, J.A., L. Neinaber, D.F. DeNardo, and A.T. Holycross. 2003. Milksnakes (Lampropeltis triangulum) from Cochise County: notes on captive breeding and pattern. Sonoran Herpetologist 16 (2):15. - get paper here
  • Baeza-Tarin, F; Hernandez, T; Giovanetto, L; Trevino, A E; Lazcano, D & Graham, S P 2018. Geographic Distribution: Lampropeltis gentilis (Western Milksnake). Herpetological Review 49 (3): 505 - get paper here
  • Baird, S. F. and C. Girard. 1853. Catalogue of North American Reptiles in the Museum of the Smithsonian Institution. Part 1.-Serpents. Smithsonian Inst., Washington, xvi + 172 pp. - get paper here
  • Barbour, Roger W. 1950. The reptiles of Big Black Mountain, Harlan County, Kentucky. Copeia 1950 (1): 100-107 - get paper here
  • Barbour, T., and L. J. Cole. 1906. Vertebrata from Yucatan. Reptilia, Amphibia and Pisces. Bull. Mus. Comp. Zool. Harvard 50: 146-159 - get paper here
  • Barts, M. 2006. Herpetologische Beobachtungen im US-Bundesstaat Oklahoma. Reptilia (Münster) 11 (61): 43-50 - get paper here
  • Bassett, L., & Pandelis, G. 2024. Updated geographic distributions for Texas reptiles. Reptiles & Amphibians, 31(1): e21492-e21492 - get paper here
  • Beane, J C 2019. New geographic distribution records for reptiles from North Carolina, USA. Herpetological Review 50 (3): 538-542 - get paper here
  • Behrendt, Marc. 2016. New herpetological county records from Perry County, Ohio, USA. Herpetological Review 47 (1): 90-91 - get paper here
  • Blair, K.B., H.M. Smith & D. Chiszar 1994. Albinism and Distributional Records for Lampropeltis triangulum (Reptilia: Serpentes) in Panhandle Texas Bull. Maryland Herpetol. Soc., 30(1): 1-5. - get paper here
  • Blanchard, Frank N. 1920. Three new snakes of the genus Lampropeltis. Occasional Papers of the Museum of Zoology, University of Michigan (81): 1-10 - get paper here
  • Blanchard, Frank N. 1920. A synopsis of the king snakes: Genus Lampropeltis Fitzinger. Occasional Papers of the Museum of Zoology, University of Michigan (87): 1-7 - get paper here
  • Blanchard, Frank N. 1928. Amphibians and reptiles of the Douglass lake region in Northern Michigan. Copeia 1928 (167): 42-51 - get paper here
  • Blatchford, D. 1989. The Jalisco Milk Snake (Lampropeltis triangulum arcifera). Litteratura Serpentium 9 (5): 234 - get paper here
  • Bocourt, M. F. 1886. In A. Duméril, M. F. Bocourt, and F. Mocquard, (1870-1909), Etudes sur les reptiles, p. i-xiv, 1-1012. In Recherches Zoologiques pour servir a l'Histoire de Ia Faune de l'Amérique Centrale et du Mexique. Mission Scientifique au Mexique et dans l'Amér Imprimerie Impériale, Paris, Vol. 3 - get paper here
  • Bossche, Johan van den 1995. Lampropeltis triangulum gentilis. Terra 31 (4): 109-114
  • Breitenbach, Luke and Erik R. Wild. 2016. Geographic Distribution: Lampropeltis triangulum (Eastern Kingsnake). Herpetological Review 47 (2): 263 - get paper here
  • BROWN, MIKALAH M. & KEITH GELUSO. 2022. Effects of cover object size and material, rainfall, and month on reptile detection in Nebraska. The Prairie Naturalist 54: 1–10. - get paper here
  • Buchert, J. 1992. Männchen von Lampropeltis triangulum sinaloae, WILLIAMS, 1978 frißt Eier des eigenen Geleges. Salamandra 28 (2): 156-157 - get paper here
  • BURBRINK, FRANK T.; JUSTIN M. BERNSTEIN, ARIANNA KUHN, MARCELO GEHARA & SARA RUANE. 2022. Ecological Divergence and the History of Gene Flow in the Nearctic Milksnakes (Lampropeltis triangulum Complex). Systematic Biology 71(4): 839–858. - get paper here
  • Burt, Charles E. 1933. A contribution to the herpetology of Kentucky. American Midland Naturalist 14 (6): 669-679 - get paper here
  • Burt, Charles E. 1935. Further records of the ecology and distribution of amphibians and reptiles in the middle west. American Midland Naturalist 16 (3): 311-336 - get paper here
  • Calderon, R.; Cedeño-Vázquez, J.R. & Pozo, C. 2003. New distributional records for amphibians and reptiles from Campeche, Mexico. Herpetological Review 34 (3): 269-272 - get paper here
  • Camper, Jeffrey D. 2019. The Reptiles of South Carolina. University of South Carolina Press, 288 pp. [review in Copeia 107 (3): 590, 2019] - get paper here
  • Canseco-Márquez, L., & Gutiérrrez-Mayén, M.G. 2010. Anfibios y reptiles del Valle de Tehuacán-Cuicatlán. Comisión Nacional para el conocimiento y uso de la biodiversidad, México D.F., Mexico, 302 pp - get paper here
  • Carrera, C. et al. 2009. Guía de Campo de los Pequeños Vertebrados del Distrito Metropolitano de Quito (DMQ). Publicación Miscelánea N° 5. Serie de Publicaciones del Museo Ecuatoriano de Ciencias Naturales (MECN) – Fondo Ambiental del MDMQ. 1-89 pp. Imprenta Nuevo Arte. Quito-Ecuador.
  • Carvajal-Cogollo, J.E.; L.E. Rojas-Murcia. & G. Cárdenas-Arévalo 2020. Reptiles del Caribe colombiano/ Reptiles of the Colombian Caribbean. Tunja: Editorial UPTC, 268 pp. - get paper here
  • Casas-Andreu, G., F.R. Méndez-De la Cruz and X. Aguilar-Miguel. 2004. Anfibios y Reptiles; pp. 375–390, in A.J.M. García-Mendoza, J. Ordoñez and M. Briones-Salas (ed.). Biodiversidad de Oaxaca. Instituto de Biología, UNAM-Fondo Oaxaqueño para la Conservación de la Naturaleza-World Wildlife Fund, México, D. F.
  • Castro-Herrera, F. & Vargas-Salinas, F. 2008. Anfibios y reptiles en el departamento del Valle del Cauca, Colombia. Biota Colombiana 9 (2): 251 - 277 - get paper here
  • Castro-Herrera, F.; Bolívar-García, W. & Herrera-Montes, M.I. 2007. Guía de los Anfibios y Reptiles des Bosque de Yotoco, Valle deel Cauca, Colombia. Universidad del Valle, Cali, Colombia, 70 pp. - get paper here
  • Chambers, E Anne; Thomas L Marshall, David M Hillis 2022. The Importance of Contact Zones for Distinguishing Interspecific from Intraspecific Geographic Variation Systematic Biology 72 (2): 357-371 [print 2023] - get paper here
  • CISNEROS-HEREDIA, D. F. 2007. On the distribution and conservation of Lampropeltis triangulum (LACÉPÈDE, 1789) in Ecuador [Short Note]. Herpetozoa 19 (3/4): 182-183 - get paper here
  • Cobarrubias, Edgar Flores; Daniel Cruz-Sáenz, and David Lazcano 2012. Notes on the Herpetofauna of Western Mexico 6: Amphibians and Reptiles of Hostotipaquillo, Jalisco, Mexico. Bull. Chicago Herp. Soc. 47 (2):21-26 - get paper here
  • Cole, Charles J.; Hardy, Laurence M. 2019. Karyotypes of six species of colubrid snakes from the Western Hemisphere, and the 140-million-year-old ancestral karyotype of Serpentes. American Museum novitates, no. 3926 - get paper here
  • Collins, J. T. ;Hirschfield, C. J. 1963. Lampropeltis doliata doliata (Linnaeus) in Kentucky. Herpetologica 19 (4): 292-293 - get paper here
  • Collins, J.T. & Collins, S.L. 2009. A pocket guide to Kansas snakes, 2nd ed. Great Plains Nature Center, Wichita, 69 pp.
  • Collins, J.T. & Collins, S.L. 2010. A pocket guide to Kansas snakes, 3rd ed. Great Plains Nature Center, Wichita, 69 pp.
  • Collins, Joseph T.;Collins, Suzanne L.;Irwin, Kelly J. 1993. Lampropeltis triangulum (milk snake). USA: Florida. Herpetological Review 24 (3): 110 - get paper here
  • Colston, Timothy J.; José António L. Barão-Nóbrega, Ryan Manders, Alice Lett, Jamie Wilmott, Gavin Cameron, Sidony Hunter, Adam Radage, Etienne Littlefair, Robert J. Williams, Antonio Lopez Cen, Kathy Slater 2015. Amphibians and reptiles of the Calakmul Biosphere Reserve, México, with new records. Check List 11 (5): 1759 - get paper here
  • Colvin, Robert. 2015. Geographic Distribution: Lampropeltis triangulum syspila (red milksnake). Herpetological Review 46 (4): 574 - get paper here
  • Conant, Roger 1938. The Reptiles of Ohio. American Midland Naturalist 20 (1): 1-200 - get paper here
  • Conant,R. & Collins,J.T. 1991. A Field Guide to Reptiles and Amphibians of Eastern/Central North America, 3rd ed. Houghton Mifflin (Boston/New York), xx + 450 p.
  • Connior, Matthew B. 2010. Geographic distribution: Lampropeltis triangulum syspila. Herpetological Review 41 (3): 380 - get paper here
  • Cope, E.D. 1860. Catalogue of the Colubridae in the Museum of the Academy of Natural Sciences of Philadelphia, with notes and descriptions of new species. Part II. Proc. Acad. Nat. Sci. Philadelphia 12: 241-266 - get paper here
  • Cope, E.D. 1889. On the snakes of Florida. Proc. US Natl. Mus. 11: 381-394 [1888] - get paper here
  • Cope, E.D. 1893. The Color Variations of the Milk Snake. American Naturalist 27 (324): 1066-1071 - get paper here
  • Cope, E.D. 1900. The crocodilians, lizards and snakes of North America. Ann. Rep. U.S. Natl. Mus. 1898: 153-1270 - get paper here
  • Cox, Christian L. and Corey E. Roelke. 2014. Geographic Distribution: Lampropeltis triangulum (milksnake). Herpetological Review 45 (3): 466 - get paper here
  • Crother, B. I. (ed.) 2012. Standard Common and Current Scientific Names for North American Amphibians, Turtles, Reptiles, and Crocodilians, Seventh Edition. Herpetological Circular 39: 1-92
  • Dallas, J., Meshaka Jr, W. E., & Delis, P. R. 2021. Evidence of Intraguild Predation in a Snake Assemblage: Bigger Seems Better. Northeastern Naturalist, 28(3), 248-260 - get paper here
  • Davis, William B.; Smith, Hobart M. 1953. Snakes of the Mexican state of Morelos. Herpetologica 8: 133-149 - get paper here
  • Degenhardt, William G.; C. W. Painter, and A. H. Price 1996. Amphibians and reptiles of New Mexico. Univ. New Mexico Press, 431 pp.
  • Dekker, Cock 2000. The escape of a Lampropeltis triangulum sinaloae. Litteratura Serpentium 20 (5): 146-147 - get paper here
  • Dixon, James R. 2000. Amphibians and reptiles of Texas, second edition. Texas A&M University Press, 421 pp.
  • Dixon, James R. and Julio A. Lemos-Espinal 2010. Amphibians and reptiles of the state of Queretaro, Mexico. Tlalnepantla UNAM, 428 pp.
  • Duellman, William E. 1947. Herpetological records from Logan County, Ohio. Copeia 1947 (3): 208 - get paper here
  • Dundee, Harold A.;Ewan, Joseph 1979. An overlooked description of a United States snake from the Eighteenth century writing of Luigi Castiglioni [Lampropeltis getulus triangulum]. Journal of Herpetology 13 (2): 216-217 - get paper here
  • Dunn, Emmett Reid 1937. Notes on tropical Lampropeltis. Occasional Papers of the Museum of Zoology, University of Michigan (353): 1-11 - get paper here
  • Duran M 2021. An annotated checklist of the amphibians and reptiles of North Padre Island, Texas, USA, with comparisons to adjacent barrier island and mainland herpetofauna. ZooKeys 1073: 119-175 - get paper here
  • Dury, R. 1932. Notes on Reptiles and Amphibians From Clifty Falls State Park, Jefferson County, Indiana. Proc. Jr. Soc. Nat. Hist. Cincinnati 3 (2): 23-26
  • Eerden, Hans van der 1992. Care and breeding of the Pueblan milk snake (Lampropeltis triangulum campbelli). Litteratura Serpentium 12 (5): 95-97 - get paper here
  • Eerden, Hans van der 1992. Care and breeding of the Pueblan milk snake (Lampropeltis triangulum campbelli). Litteratura Serpentium 12 (3): 52-53 - get paper here
  • Elmberg, J. 1980. Herpetological observations 77-78 [in Vernon county, Wisconsin]. Fältbiologerna Umea.
  • Enderson, Erik F.; Thomas R. Van Devender, Robert L. Bezy 2014. Amphibians and reptiles of Yécora, Sonora and the Madrean Tropical Zone of the Sierra Madre Occidental in northwestern Mexico. Check List 10 (4): 913-926 - get paper here
  • Enge, Kevin M. 2009. Venomous and non-venomous snakes of Florida. Publication of the Florida Fish & Wildlife Conservation Commission. 16 pp.
  • Ernst, C.H., & Barbour, R.W. 1989. Snakes of eastern North America. George Mason Univ. Press, Fairfax, VA 282 pp.
  • Fitch, H.S. & R.R. Fleet 1970. Natural History of the milk snake (Lampropeltis triangulum) in northeastern Kansas Herpetologica 26 (4): 387-396. - get paper here
  • Foster, N. & Conway, C. 2012. Geographic distribution: Lampropeltis triangulum (milksnake). Herpetological Review 43: 449 - get paper here
  • Fowler, Henry W. 1915. Some amphibians and reptiles of Cecil Country, Maryland [Lampropeltis triangulum doliata]. Copeia 1915 (22): 37-40 - get paper here
  • FRANZ, L. R. 1966. An Abnormal Pattern in the Eastern Milk Snake, Lampropeltis doliata triangulum (Lacepede). Bull. Maryland Herp. Soc. 2 (1): 11 - get paper here
  • Gabirot, Marianne; Pablo Picerno, Jorge Valencia, Pilar Lopez, and José Martin 2012. Species Recognition by Chemical Cues in Neotropical Snakes. Copeia 2012 (3): 472-477. - get paper here
  • García, A. & Ceballos, G. 1994. GUIA DE CAMPO DE LOS REPTILES Y ANFIBIOS DE LA COSTA DE JALISCO, MEXICO. FUNDACION ECOLOGICA DE CUIXMALA, A.C. INSTITUTO DE BIOLOGIA, UNAM - get paper here
  • Garman, S. 1887. Reptiles and batrachians from Texas and Mexico. Bull. Essex Inst. 19: 119-138
  • Garman, Samuel 1884. The reptiles and batrachians of North America. Mem. Mus. comp. Zool, Cambridge (Massachusetts), 8 (3): xxxiv + 185 pp. [1883] [CNAH reprint 10] - get paper here
  • Gehlbach, Frederick R. 1962. Aberrant western milksnake, Lampropeltis doliata Linnaeus, from New Mexico. Southwestern Naturalist 7 (3/4): 270-272 - get paper here
  • GLORIOSO, BRAD. 2021. New amphibian and reptile parish records from Louisiana, USA. Herpetological Review 52 (2): 364–366.
  • Gray, Brian S. 2011. Seasonal Activity and Natural History Observations of Five Snake Species from the Central Lowland Province of Erie County, Pennsylvania. Journal of Kansas Herpetology (38): 13 - get paper here
  • Green, N.B., & Pauley, T.K. 1987. Amphibians and reptiles in West Virginia. Univ. of Pittsburgh Press, Pittsburgh, 241 pp.
  • Greene, H.W. 1979. (Book review) Systematics and natural history of the American milksnake, Lampropeltis triangulum by K.L. Williams Copeia 1979 (1): 180-181. - get paper here
  • Greene, Harry W. 1979. Systematics and natural history of the American milksnake, Lampropeltis triangulum. Copeia 1979 (1): 180-181 - get paper here
  • Greene, Harry W. 1981. Literature: (Review of) Systematics and natural history of the American milk snake, Lampropeltis triangulum. Litteratura Serpentium 1 (5): 206 - get paper here
  • Gregory, P.T. 2004. Analysis of patterns of aggregation under cover objects in an assemblage of six species of snakes. Herpetologica 60 (2): 178-186 - get paper here
  • Gregory, Patrick T. 2015. Natural history notes on a northern population of the milksnake (Lampropeltis triangulum), with evidence for ontogenetic shifts in cover use and diet. Herpetological Review 46 (4): 506-509 - get paper here
  • Grogan, W. L., Jr. and D. C. Forester 1998. New records of the milk snake, Lampropeltis triangulum, from the Coastal Plain of the Delmarva Peninsula, with comments on the status of L. t. temporalis. The Maryland Naturalist 42:5-14.
  • Gross, Iwo P., Wang, Yong and Schweitzer, Callie J. 2016. Lampropeltis triangulum syspila (Red Milksnake) Piebaldism. Herpetological Review 47 (4): 684-685 - get paper here
  • Groves, John D. and Peter S. Sachs 1973. Eggs and Young of the Scarlet King Snake, Lampropeltis triangulum elapsoides. Journal of Herpetology 7 (4): 389-390 - get paper here
  • GUTIÉRREZ MAYÉN, MA. GUADALUPE Y JORGE SALAZAR ARENAS 2007. HERPETOFAUNA DE LOS MUNICIPIOS DE CAMOCUAUTLA, ZAPOTITLÁN DE MÉNDEZ Y HUITZILAN DE SERDÁN, DE LA SIERRA NORTE DE PUEBLA. HERPETOFAUNA DE TRES MUNICIPIOS DE LA SIERRA NORTE DE PUEBLA, pp.197-223
  • Gutsche, A. 2012. Klassiker: die Dreiecksnatter. Terraria-Elaphe 2012 (5): 80-81 - get paper here
  • Guyer, Craig; Mark A. Bailey, and Robert H. Mount 2018. Lizards and snakes of Alabama. University of Alabama Press, 397 pp. - get paper here
  • Hall, Charles W. 1951. Notes on a small herpetological collection from Guerreo. Univ. Kansas Sci. Bull. 34 (4): 201-212 - get paper here
  • Hallmen, M. 2005. Farb- und Zeichnungszuchten in der Terraristik. Reptilia (Münster) 10 (55): 16-22 - get paper here
  • Hallmen, M. 2006. Selective Breeding for color and pattern. Reptilia (GB) (44): 12-18 - get paper here
  • Hamilton, Bryan Tyler; Rachel Hart, and Jack W. Sites 2012. Feeding Ecology of the Milksnake (Lampropeltis triangulum, Colubridae) in the Western United States. Journal of Herpetology 46 (4): 515-522. - get paper here
  • Harris, H.S., Jr. 2006. Partial Albinism in Maryland Lampropeltis triangulum temporalis Bull. Maryland Herpetol. Soc., 42(4): 190-192. - get paper here
  • Hedges, S. B. 1977. The presence of the scarlet kingsnake, Lampropeltis triangulum elapsoides Hollbrook (Reptilia, Serpentes, Colubridae), in the Florida Keys. Herpetological Review 8 (4): 125-126 - get paper here
  • Herman, Dennis W. 1980. Literature: Breeding the Jaliscan milk snake (Lampropeltis triangulum arcifera) at Atlanta Zoo. Litteratura Serpentium 1 (1): 31 - get paper here
  • HERTZ, A. & VAN DEN BERGHE, E. 2009. Atypical coloration pattern in a specimen of Lampropeltis triangulum stuarti WILLIAMS, 1978, from western Nicaragua [Short Note]. Herpetozoa 22: - get paper here
  • Hileman, Eric T.; Joshua M. Kapfer, Timothy C. Muehlfeld, and John H. Giovanni 2015. Recouping Lost Information when Mark-Recapture Data are Pooled: A Case Study of Milksnakes (Lampropeltis triangulum) in the Upper Midwestern United States. Journal of Herpetology Sep 2015, Vol. 49, No. 3: 428-436. - get paper here
  • Hillis, D. M., Chambers, E. A., & Devitt, T. J. 2021. Contemporary Methods and Evidence for Species Delimitation. Ichthyology & Herpetology, 109(3), 895-903 - get paper here
  • Hinds, Brian. 2010. Geographic distribution: Lampropeltis triangulum gentilis. Herpetological Review 41 (3): 380 - get paper here
  • Hingley, Kevin 1982. Literature: Notes on keeping the Scarlet king snake (Lampropeltis triangulum elapsoides). Litteratura Serpentium 2 (6): 308-309 - get paper here
  • Holt, B. D. and C. Rose 2016. Geographic Distribution: Lampropeltis triangulum (Milksnake). Herpetological Review 47(3): 427. - get paper here
  • HOLT, B. D., AND Z. VEAL 2020. Geographic Distribution: Lampropeltis triangulum (Eastern Milksnake). USA: Alabama: Cherokee Co. Herpetological Review 51: 80.
  • Holycross, Andrew T.;Simonson, Rob 1998. Geographic Distribution. Lampropeltis triangulum. Herpetological Review 29 (2): 113-114 - get paper here
  • Hopkins, Terry. 2014. Geographic Distribution: Lampropeltis triangulum (millksnake). Herpetological Review 45 (1): 95 - get paper here
  • Howard, C.J. 1989. Husbandry and Breeding of the Honduran Milk Snake (Lampropeltis triangulum hondurensis) at Twycross Zoo. Litteratura Serpentium 9 (5): 233-234 - get paper here
  • Hromada, S J; Snyder, S D & Gienger, C M; 2018. Geographic Distribution: Lampropeltis triangulum (Eastern Milksnake). Herpetological Review 49 (4): 718 - get paper here
  • Hubbs, B. 2012. Geographic distribution: Lampropeltis triangulum syspila (red milk snake). Herpetological Review 43: 621 - get paper here
  • Hubbs, Brian 1998. Geographic Distribution. Lampropeltis triangulum celaenops. Herpetological Review 29 (1): 54 - get paper here
  • Hudson, Bryan D., Zach I. Felix, Justin Oguni, Brad Wilson, Kira McEntire, Theresa Stratmann, Daniel D. Duff and Zack Seymour. 2015. New geographic distributional records of amphbians and reptiles in Georgia, USA. Herpetological Review 46 (4): 595-596 - get paper here
  • Hunter, M.L., Jr., Albright, J., & Arbuckle, J., eds. 1992. The amphibians and reptiles of Maine. Maine Agric. Experiment Station, Orono, Bull. 838 188 pp.
  • Husak, Jerry F.;Wright, Jeff 1998. Geographic Distribution. Lampropeltis triangulum. Herpetological Review 29 (2): 114 - get paper here
  • Hyatt, D.M. 1989. Systematics and Natural History of the American Milk Snake, Lampropeltis triangulum (Book Review). Tucson Herpetological Society Newsletter 2(4):34-35.
  • ICZN 1987. Opinion 1463. De Lacepede 1788-1789, Histoire Naturelle des Serpens and later editions: rejected as a non-binominal work. Bull. Zool. Nomenclature 44(4): 265-267. - get paper here
  • Irwin, Kelly J. 2004. Arkansas Snake Guide. Arkansas Game & Fish Commission, Little Rock, 50 pp.
  • Irwin, Kelly J.;Collins, Suzanne L. 1996. Geographic Distribution. Lampropeltis triangulum. Herpetological Review 27 (1): 34 - get paper here
  • Jan, G. 1865. Iconographie générale des ophidiens. 14. Livraison. [Elapomorphus d’Orbignyi, Coronella getulus var. Sayi]. J.B. Bailière et Fils, Paris - get paper here
  • Jan, G. 1866. Iconographie générale des ophidiens. 17. Livraison. J.B. Bailière et Fils, Paris - get paper here
  • Jensen, John B.; Carlos D. Camp, Whit Gibbons, & Matt J. Elliott 2008. Amphibians and reptiles of Georgia. University of Georgia Press, 575 pp.
  • Johnson, H. 2017. Geographic Distribution: Lampropeltis gentilis (Western Milksnake). Herpetological Review 48 (2): 391 - get paper here
  • Kapfer, Joshua M., Jeffrey M. Lorch, Erik R. Wild, Donald J. Brown, Lisa Mitchem, Nicholas Rudolph, Karl Rutzen and Richard C. Vogt. 2015. Distributional records for amphibians and reptiles from Wisconsin, USA. Herpetological Review 46 (4): 587-590 - get paper here
  • Kardon, Alan 1980. Literature: A note on captive reproduction in three Mexican milk snakes (Lampropeltis triangulum polyzona, L.t. nelsoni and L.t. sinaloae). Litteratura Serpentium 1 (1): 31 - get paper here
  • Kennicott, R. 1860. Descriptions of new species of North American serpents in the museum of the Smithsonian Institution, Washington. Proc. Acad. Nat. Sci. Philadelphia 12: 328-338 - get paper here
  • Keogh, J. Scott 1996. Evolution of the colubrid snake tribe Lampropeltini: a morphological perspective. Herpetologica 52 (3): 406-416 - get paper here
  • Köhler, G. 2008. Reptiles of Central America. 2nd Ed. Herpeton-Verlag, 400 pp.
  • Köhler, G., J. R. Cedeño-Vázquez, E. D. Kraus, P. M. Beutelspacher-García, and J. A. Domínguez-Lepe. 2017. The Chetumal Snake Census: generating biological data from road-killed snakes. Part 5. Imantodes tenuissimus, Lampropeltis triangulum, and Stenorrhina freminvillii. Mesoamerican Herpetology 4(4): 774–789 - get paper here
  • Kornacker,P.M. 1999. Checklist and key to the snakes of Venezuela. PaKo-Verlag, Rheinbach, Germany, 270 pp.
  • Kraus, Fred; Schuett, Gordon W. 1981. An integrade Lampropeltis triangulum from West Texas. Herpetological Review 12 (2): 53 - get paper here
  • Kreutz, R. 2005. Farb- und Zeichnungsstandard der Kornnatter (Panterhophis guttatus). Kirschner und Seufer Verlag, Keltern-Weiler, 158 pp.
  • Kuch, U. 1988. Das Portrait: Lampropeltis triangulum micropholis COPE. Sauria 10 (4): 1-2 - get paper here
  • Kwet, Axel 2015. Liste der im Jahr 2014 neu beschriebenen Reptilien. Terraria-Elaphe 2015 (3): 50-64 - get paper here
  • Lacepède, B. G. E. 1789. Histoire Naturelle des Quadrupèdes Ovipares et de Serpens. Vol.2. lmprimerie du Roi, Hôtel de Thou, Paris, 671 pp. - get paper here
  • LAFLEUR, RICHARD M. 2022. Geographic distribution: LAMPROPELTIS TRIANGULUM (Eastern Milksnake). USA: WISCONSIN: Clark Co. Herpetological Review 53 (1): 79.
  • Laita, Mark 2013. Serpentine. Abrams and PQ Blackwell, Auckland, New Zealand, 200 unnumbered pages
  • Langner, Christian 2014. Herpetologische Beobachtungen im Lebensraum von Abronia deppii im Valle de Bravo, Estado de México, Mexiko. Terraria-Elaphe 2014 (2): 46-55 - get paper here
  • Largelamb, H. A. 1907. Notes on the remarkable habits of certain turtles and lizards. National Geographic 18(6):413-419
  • Lazcano D, Nevárez-de los Reyes M, García-Padilla E, Johnson JD, Mata-Silva V, DeSantis DL, Wilson LD. 2019. The herpetofauna of Coahuila, Mexico: composition, distribution, and conservation status. Amphibian & Reptile Conservation 13(2) [General Section]: 31–94 (e189) - get paper here
  • LeClere, Jeff 1994. Snakes of Minnesota: Eastern milksnake (Lampropeltis triangulum triangulum). Newsletter of the Minnesota Herpetological Society 14 (12): 15-17
  • Lee, J. C. 2000. A field guide to the amphibians and reptiles of the Maya world. Cornell University Press, Ithaca,
  • Lee, J.C. 1996. The amphibians and reptiles of the Yucatán Peninsula. Comstock, Cornell University Press, Ithaca, 500 pp.
  • Lemos-Espinal JA, Smith GR 2016. Amphibians and reptiles of the state of Coahuila, Mexico, with comparison with adjoining states. ZooKeys 593: 117-137, doi: 10.3897/zookeys.593.8484 - get paper here
  • Lemos-Espinal JA, Smith GR, Cruz A 2016. Amphibians and Reptiles of the state of Nuevo León, Mexico. ZooKeys 594: 123-141, doi: 10.3897/zookeys.594.8289 - get paper here
  • Lemos-Espinal, Julio A. and James R. Dixon 2013. Amphibians and Reptiles of San Luis Potosí. Eagle Mountain Publishing, xii + 300 pp.
  • Lillywhite, Harvey B. 2014. How Snakes Work: Structure, Function and Behavior of the World's Snakes. Oxford University Press, New York, 256 pp
  • Linzey, D.W., & Clifford, M.J. 1981. Snakes of Virgina. Univ. Press of Virginia, Charlottesville 159 pp.
  • LIttle, Raymond N. 2010. Geographic distribution: Lampropeltis triangulum. Herpetological Review 41 (3): 380 - get paper here
  • Loomis, Richard B. 1949. The red milk snake, Lampropeltis doliata syspila, in Nebraska. Copeia 1949 (4): 295 - get paper here
  • Love, B. 2012. Amerikanische Herp-Probleme. Reptilia (Münster) 17 (98): 18-19 - get paper here
  • Lowe, C.H. 1989. Arizona's rarest snake - Lampropeltis triangulum, the milksnake. Tucson Herpetological Society Newsletter 2(1):5.
  • Luja VH, López JA, Cruz-Elizalde R, Ramírez-Bautista A 2017. Herpetofauna inside and outside from a natural protected area: the case of Reserva Estatal de la Biósfera Sierra San Juan, Nayarit, Mexico. Nature Conservation 21: 15-38 - get paper here
  • Maddalena, Marcus P.; Jeffrey R. Row, Matthew E. Dyson, Gabriel Blouin-Demers, Bradley C. Fedy 2020. Movement and Habitat Selection of Eastern Milksnakes (Lampropeltis triangulum) at Intact and Fragmented Sites. Copeia 108 (4): 847-854 - get paper here
  • Marcou, Zachary John 2018. Delineating Metrics of Diversity for a Snake Community in a Rare Ecosystem. MSc Thesis, Stephen F. Austin State University - get paper here
  • Martof, Bernard 1955. Some reptiles of Georgia. Copeia 1955 (4): 302-305 - get paper here
  • Mattison, Chris 2007. The New Encyclopedia of Snakes. Princeton University Press
  • McAllister, C. T. & Robison, H. W. 2012. Geographic distribution: Lampropeltis triangulum syspila (milksnake). Herpetological Review 43: 449 - get paper here
  • McAtee, W.L. 1907. A list of the mammals, reptiles and batrachians of Monroe County, Indiana. Proc. Biol. Soc. Washington 20: 1-16 - get paper here
  • McCauley, Robert H., Jr. 1941. A redescription of Lampropeltis triangulum temporalis (Cope). Copeia 1941 (3): 146-150 - get paper here
  • McCranie J R 2011. The snakes of Honduras. SSAR, Salt Lake City, 725 pp.
  • McCranie, J. & Castañeda, F.E. 2005. The herpetofauna of Parque Nacional Pico Bonito, Honduras. Phyllomedusa 4 (1): 3-16 - get paper here
  • McCrystal, Hugh K.;dean, Robert H.;Dixon, James R. 1984. Lampropeltis triangulum annulata (Mexican Milk Snake). Size. Herpetological Review 15 (1): 19 - get paper here
  • Mehrtens, J.M. 1987. Living snakes of the world in color. Sterling Publ. Co., hic., New York, NY: 480 pp.
  • Mense, Mark 2010. Welche Schlange für mich? Reptilia (Münster) 15 (86): 51-56 - get paper here
  • Mertens, R. 1952. Die Amphibien und Reptilien von El Salvador. Abh. senckenb. naturf. Ges. (Frankfurt) (No. 487): 120 pp.
  • Meshaka Jr, Walter E.;, Jane Netting Huff & Robert C. Leberman 2008. Amphibians and Reptiles of Powdermill Nature Reserve in Western Pennsylvania. Journal of Kansas Herpetology (25): 12-18 - get paper here
  • Meshaka, Jr., Walter E. & Joseph T. Collins 2009. A POCKET GUIDE TO PENNSYLVANIA SNAKES. The Commonwealth of Pennsylvania, 56 pp. - get paper here
  • Miller, R. & G. Grall 1978. Reproductive Data on Lampropeltis triangulum temporalis from Maryland Bull. Maryland Herpetol. Soc., 14(1): 36-38. - get paper here
  • Mitchell, J. C. & Reay, K.K. 1999. Atlas of amphibians and reptiles in Virginia. Specialty Publication 1, VA Dept of Game and Fisheries, 122 pp. - get paper here
  • Mitchell, J.C. 1994. The reptiles of Virginia. Virginia Department of Game and Inland Fisheries, ca. 350 pp.
  • Moerkens, Tony 1993. My breeding of Lampropeltis triangulum sinaloae. Litteratura Serpentium 13 (3): 83-88 - get paper here
  • Montgomery, Chad; Childers, Theresa; Bergman, Enoch; Manzer, Jerry D.; Sifert, James; Hill, Ben; Mackessy, Stephen P. 1998. Geographic Distribution. Lampropeltis triangulum gentilis. Herpetological Review 29 (2): 114 - get paper here
  • Montgomery, Chad;Hobert, Justin;Bergman, Enoch;Hill, Ben;Boback, Scott;Mackessy, Stephen P. 1997. Geographic Distribution. Lampropeltis triangulum gentilis. Herpetological Review 28 (1): 52 - get paper here
  • Muell, Morgan R. 2016. Geographic Distribution: Lampropeltis triangulum triangulum (Eastern Milksnake). Herpetological Review 47 (4): 630 - get paper here
  • Mulaik, Stanley; Mulaik, Dorthea 1945. Lampropeltis triangulum annulata from Kerr County, Texas. Copeia 1945 (1): 49 - get paper here
  • Muñoz-Alonso, Luis Antonio; Jorge Nieblas-Camacho,<br>Marina Alba Chau-Cortez, Alondra Berenice González-Navarro, Jaime López-Pérez & Juan Pérez-López 2017. Diversidad de anfibios y reptiles en la Reservade la Biosfera Selva El Ocote: su vulnerabilidad ante la fragmentación y el cambio climático. In: Lorena Ruiz-Montoya et al. (eds), Vulnerabilidad social y biológica ante el cambio climático en la Re El Colegio de la Frontera Sur, pp.395-448 - get paper here
  • Nelson, D.J. 1948. Lampropeltis triangulum gentilis in Montana Herpetologica 4 (5): 170. - get paper here
  • Nelson, Dee Jay 1950. Lampropeltis triangulum gentilis in Montana. Herpetologica 6: 41-44 - get paper here
  • Nicholson, Abbigail E. 2013. Geographic Distribution: Lampropeltis triangulum (milksnake). Herpetological Review 44 (3): 477 - get paper here
  • Nordgren, Matt and Chad Whitney 2011. Lampropeltis triangulum (Milk Snake) Kansas. Journal of Kansas Herpetology (38): 7 - get paper here
  • Norrie, S.G. 1989. Comparisons between the natural and captive environments of the Sinaloan milk snake (Lampropeltis triangulum sinaloae). Litteratura Serpentium 9 (6): 284 - get paper here
  • O’Shea, M. 2018. The Book of Snakes. Ivy Press / Quarto Publishing, London, - get paper here
  • Páez, V.P. et al. 2002. Guía de campo de algunas especies de anfibios y reptiles de Antioquia. Universidad de Antioquia, - get paper here
  • Painter, Charles W.; Jennings, Randy D. 1996. Geographic Distribution. Lampropeltis triangulum celaenops. Herpetological Review 27 (4): 213 - get paper here
  • Palacio-Baena, Jaime Alberto; Eliana María Muñoz Escobar; Sandra Milena Gallo Delgado; Mauricio Rivera Correa 2006. ANFIBIOS Y REPTILES DEL VALLE DE ABURRÁ Editorial Zuluaga, 92 pp. - get paper here
  • Palmer, W.M. & Braswell, A.L. 1995. Reptiles of North Carolina. Univ. North Carolina Press
  • Parmley, Dennis 1994. Reevaluation of the extinct kingsnake Lampropeltis intermedius Brattstrom (Colubridae) with comments on the ancestry of Lampropeltis triangulum. Herpetological Natural History 2 (2): 83-88
  • Paterson, J. E., Pulfer, T., Horrigan, E., Sukumar, S., Vezina, B. I., Zimmerling, R., & Davy, C. M. 2021. Individual and synergistic effects of habitat loss and roads on reptile occupancy. Global Ecology and Conservation, 31, e01865 - get paper here
  • PEALE, M., B. LONG, AND B. L. CHRISTMAN 2020. Geographic Distribution: Lampropeltis gentilis (Western Milksnake). USA: New Mexico: Taos Co. Herpetological Review 51: 79-80.
  • Pérez-Santos,C. & Moreno, A.G. 1988. Ofidios de Colombia. Museo reegionale di Scienze Naturali, Torino, Monographie VI, 517 pp.
  • Peterson, Karl H., David Lazcano & Ramiro D. Jacobo Galván 1995. Captive reproduction in the Mexican milksnake Lampropeltis triangulum annulata. Litteratura Serpentium 15 (5): 128-132 - get paper here
  • Phillips, JG 2016. Updated Geographic Distributions of Michigan Herpetofauna: a Synthesis of Old and New Sources. The Journal of North American Herpetology 2016: 45–69 - get paper here
  • Plummer, Michael V. and Steven E. Seitz. 2016. Geographic Distribution: Lampropeltis triangulum amaura (Louisiana Milksnake). Herpetological Review 47 (2): 263 - get paper here
  • Porras, L.W. 2006. Die Schlangen des costa-ricanischen Trockenwaldes. Reptilia (Münster) 11 (61): 32-37 - get paper here
  • Porras, L.W. & Solórzano, A. 2006. Costa Rica’s venomous snakes. Reptilia (GB) (48): 11-17 - get paper here
  • Porras, Louis 1996. Exquiaite!!! A kaleidoscope of colors: Honduran milks snakes Lampropeltis triangulum hondurensis. Vivarium 7 (5): 28-37
  • Powell, R. & Parmerlee, J. 1981. Eine bisher unbeschriebene Farbvariante von Lampropeltis triangulum blanchardi (Reptilia: Serpentes: Colubridae). Salamandra 17 (1-2): 83-86 - get paper here
  • Price, Andrew H.;Johnson, Dane G. 1978. Lampropeltis triangulum celaenops (New Mexico Milk Snake). USA: New Mexico. Herpetological Review 9 (3): 108 - get paper here
  • Pyron, R.Alexander; Frank T. Burbrink 2009. Neogene diversification and taxonomic stability in the snake tribe Lampropeltini (Serpentes: Colubridae). Molecular Phylogenetics and Evolution 52 (2): 524-529 - get paper here
  • Quinn, H. R. 1983. Two new subspecies of Lampropeltis triangulum from México. Trans. Kansas Acad. Sci. 86: 113-135. - get paper here
  • Rapier, B C; Smith, L P; Hromada, S J & Gienger, C M; 2019. Geographic Distribution: Lampropeltis triangulum (Eastern Milksnake) USA: Tennessee: Cumberland Co. Herpetological Review 50 (1): 106 - get paper here
  • Repp, R., J. Feldner, R. Babb, R.B. Duncan, A.T. Holycross, E.M. Nowak, and M. Demlong. 1999. Arizona milk snake; an e-mail debate. Sonoran Herpetologist 12 (10):108-111. - get paper here
  • Riel, C.A.P. van 1983. Lampropeltis triangulum elapsoides, the Scarlet kingsnake. Litteratura Serpentium 3 (6): 193-202 - get paper here
  • RIVAS, GILSON A.; CÉSAR R. MOLINA, GABRIEL N. UGUETO, TITO R. BARROS, CÉSAR L. BAR- RIO-AMORÓS & PHILIPPE J. R. KOK 2012. Reptiles of Venezuela: an updated and commented checklist. Zootaxa 3211: 1–64 - get paper here
  • Rodríguez, M.C. & H. Drummond 2000. Exploitation of Avian Nestlings and Lizards by Insular Milksnakes, Lampropeltis triangulum Journal of Herpetology 34 (1): 139-142. - get paper here
  • Rojas-Morales, Julián Andrés 2012. Snakes of an urban-rural landscape in the central Andes of Colombia: species composition, distribution, and natural history. Phyllomedusa 11 (2): 135- - get paper here
  • Roth, J.J.;Smith, H.M. 1990. The milksnake, Lampropeltis triangulum, in northwest Colorado. Bull. Chicago Herp. Soc. 25 (1): 6-7 - get paper here
  • Ruane, Sara; Robert W. Bryson, Jr., R. Alexander Pyron, and Frank T. Burbrink 2014. Coalescent Species Delimitation in Milksnakes (genus Lampropeltis) and Impacts on Phylogenetic Comparative Analyses. Systematic Biology 63 (2): 231-250, doi:10.1093/sysbio/syt099 - get paper here
  • Savage, J.M. 2002. The Amphibians and Reptiles of Costa Rica: A Herpetofauna Between Two Continents, Between Two Seas. University of Chicago Press, 934 pp. [review in Copeia 2003 (1): 205]
  • Schaefer, Kurt;Chiszar, David;Smith, Hobart M. 1995. Lampropeltis triangulum gentilis. Herpetological Review 26 (2): 110 - get paper here
  • Schmidt, D. 2004. Zur Reproduktion von Schlangen im Terrarium: Aufgaben - Fakten - Probleme. Draco 5 (17): 22-36 - get paper here
  • Schmidt, D. 2015. Nattern. Arten und Tipps für Einsteiger. Reptilia (Münster) 20 (115): 28-41 - get paper here
  • Schmidt, D. & Kunz, K. 2005. Ernährung von Schlangen. Natur und Tier Verlag, Münster, 159 pp. - get paper here
  • Schmidt, Dieter 2006. Die Dreiecksnatter (Lampropeltis triangulum). Natur und Tier Verlag (Münster), 64 pp. - get paper here
  • Schmidt, T. 2006. Haltung und Nachzucht der Puebla-Königsnatter, Lampropeltis triangulum campelli Terraria (2) 1 (2): 59-62.
  • Schmidt, T. 2008. Lampropeltis triangulum andesiana - seltenes Terrarien-Juwel mit zweifelhaftem Ruf. Teil 2: Vermehrung Terraria (14) 3 (6): 46-49.
  • Schmidt, T. 2008. Lampropeltis triangulum andesiana - seltenes Terrarien-Juwel mit zweifelhaftem Ruf. Teil 1: Biologie und Haltung Terraria (13) 3 (5): 54-59.
  • Schmidt, T. 2011. Haltung und Nachzucht der Pazifik-Dreiecksnatter, Lampropeltis triangulum oligozona. Reptilia (Münster) 16 (91): 52-57 - get paper here
  • Schuett, Gordon W. 1994. Lampropeltis triangulum (milk snake). USA: Wyoming. Herpetological Review 25 (4): 167 - get paper here
  • Shaw, George & Nodder, Frederick P. (Eds.) 1796. The Naturalist's Miscellany [...], Vol. VII. London, Nodder & Co., plates 219-254, 136 unnumbered pages [published in monthly issues between August 1, 1795, and July 1, 1796] - get paper here
  • Shine, Richard 1994. Sexual size dimorphism in snakes revisited. Copeia 1994 (2): 326-346 - get paper here
  • Shreve, B. 1936. An Unusual Habitat Record for Lampropeltis triangulum gentilis. Copeia, 1936(1), 72-72 - get paper here
  • Smith, H.M., K.L. Williams & D. Chiszar 2000. The Prepublication History of the Name Lampropeltis triangulum sinaloae Williams, 1978, for the Sinaloan Milk Snake Bull. Maryland Herpetol. Soc., 36(1): 5-8. - get paper here
  • Smith, Hobart M. 1942. Remarks on the Mexican king snakes of the triangulum group. Proc. Rochester Academy of Science 8 (4): 196-207 - get paper here
  • Smith, Hobart M.;Ristau, Carolyn;Bell, Tom;Bell, Marion;Chiszar, David 1995. Lampropeltis triangulum multistriata x taylori. Herpetological Review 26 (4): 210 - get paper here
  • Smith, J E & MIller, B T; 2019. Geographic Distribution: Lampropeltis triangulum (Milksnake) USA: Tennessee: Lawrence Co. Herpetological Review 50 (3): 529 - get paper here
  • Smith, Philip W. 1956. A Blotch-Count Gradient in Snakes. Herpetologica 12: 156-160 - get paper here
  • Spinner, Leo 2017. Auf Schlangensuche in Cape Cod, Massachusetts. Terraria-Elaphe 2017 (1): 14-24 - get paper here
  • Spuckler, T. & Durson, A.M. 2017. Lampropeltis triangulum (Milksnake) Predation. Herpetological Review 48 (4): 859-860 - get paper here
  • Stebbins,R.C. 1985. A Field Guide to Western Reptiles and Amphibians, 2nd ed. Houghton Mifflin, Boston
  • Stejneger, L. 1891. Notes on some North American snakes. Proc. US Natl. Mus. 14 (876): 501-505 - get paper here
  • Stejneger, L. 1902. The reptiles of the Huachuca Mountains, Arizona. Proc. US Natl. Mus. 25 [1902]: 149-158 - get paper here
  • Stejneger, L.H., and T. Barbour 1917. A Check List of North American Amphibians and Reptiles. Harvard University Press, Cambridge. 4: i-iv, 1-125 - get paper here
  • STEVENSON, DIRK J. et al. 2021. New County Records for Amphibians and Reptiles in Georgia, USA. Herpetological Review 52 (2): 350-359
  • Strimple, Pete 1991. Captive reproduction in the Scarlet kingsnake, Lampropeltis triangulum elapsoides (Holbrook, 1838). Litteratura Serpentium 11 (3): 50-52 - get paper here
  • Stuart, L. C. 1935. A contribution to a knowledge of the herpetology of a portion of the savanna region of Central Peten, Guatemala. University of Michigan Museum of Zoology Miscellaneous Publications 29: 1-56 - get paper here
  • Stuart, L. C. 1935. Studies on neotropical colubrinae. V. A new snake of the genus Lampropeltis from Yucatan. Occasional Papers of the Museum of Zoology, University of Michigan (309): 1-6 - get paper here
  • Tanner, W.W. Loomis, R.B. 1957. A taxonomic and distributional study of the western subspecies of the milksnake, Lampropeltis doliata. Transactions of the Kansas Academy of Science 60 (1): 12-42 - get paper here
  • Taylor,E.H. 1949. A preliminary account of the herpetology of the state of San Luis Potosi, Mexico. Univ. Kansas Sci. Bull. 33 (2): 169-215 - get paper here
  • Tennant, A. 2003. Snakes of North America - Eastern and Central Regions, revised edition. Lone Star Books, 605 pp.
  • Tennant, A. & Bartlett, R.D. 2000. Snakes of North America - Eastern and Central Regions. Gulf Publishing, Houston, TX, 588 pp.
  • Terrell, Vanessa C., Jaimie L. Klemish, Nathan J. Engbecht, John A. May, Peter J. Lannoo, Rochelle M. Stiles, and Michael J. Lannoo. 2014. Amphibian and reptile colonization of reclaimed coal spoil grasslands. The Journal of North American Herpetology 2014(1):59-68 - get paper here
  • Thissen, R. & Hansen, H. 2001. Königsnattern - Lampropeltis. Natur und Tier Verlag (Münster), 172 pp. - get paper here
  • Torre-Loranca, M.A. de la; Aguirre-León, G. & López-Luna, M.A. 2006. Coralillos verdaderos (Serpentes: Elapidae) y Coralillos falsos (Serpentes: Colubridae) de Veracruz, México. Acta Zoologica Mexicana 22 (3): 11-22 - get paper here
  • TREFRY, I. W., AND J. G. HIMES 2020. Geographic Distribution: Lampropeltis gentilis (Western Milksnake). USA: Louisiana: Webster Parish. Herpetological Review 51: 546.
  • TRYON, B. W., & MURPHY, J. B. 1982. Miscellaneous Notes on the Reproductive Biology of Reptiles. 5. Thirteen Varieties of the Genus Lampropeltis, species mexicana, triangulum and zonata. Trans. Kansas Acad. Sci. 85 (2): 96-119 - get paper here
  • Valencia-Zuleta A, Jaramillo-Martínez AF, Echeverry-Bocanegra A, Viáfara-Vega R, Hernández-Córdoba O, Cardona-Botero VE, Gutiérrez-Zúñiga J, Castro-Herrera F. 2014. Conservation status of the herpetofauna, protected areas, and current problems in Valle del Cauca, Colombia. Amphibian & Reptile Conservation 8 (2): 1–18 (e87) - get paper here
  • Vance, Thomas 1994. Lampropeltis triangulum amaura (Louisana milk snake). USA: Texas. Herpetological Review 25 (2): 77 - get paper here
  • VanDeWalle, Terry 2022. The Natural History of the Snakes and Lizards of Iowa. University of Iowa Press, Iowa City, Iowa, 384 pp. ISBN: 978-1- 609388-37-9. [review in HR 54 (1): 151] - get paper here
  • Wallach, Van 2005. A new dicephalic Lampropeltis t. triangulum from Maine, USA Podarcis 6 (1/2): 36-39 - get paper here
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp.
  • Walley, Harlad D. 1997. Geographic Distribution. Lampropeltis triangulum triangulum. Herpetological Review 28 (3): 159 - get paper here
  • Walley, Harlan D.; Queral-Regil, Alejandro; Bittner, Tonya 1998. Geographic Distribution. Lampropeltis triangulum triangulum. Herpetological Review 29 (1): 54 - get paper here
  • Wanrooy, V.J. van 1999. Breeding results: Lampropeltis triangulum campbelli. Litteratura Serpentium 19 (2): 45 - get paper here
  • Wanrooy, V.J. van 1999. Breeding results: Lampropeltis triangulum sinaloae. Litteratura Serpentium 19 (2): 45 - get paper here
  • Webb, R.G. 1984. Herpetogeography in the Mazatlán-Durango Region of the Sierra Madre Occidental, Mexico. Vetrebrate Ecology and Systematics - A ribute to Henry S. Fitch; Museum of Natural History, University of Kansas, Lawrence, pp. 217-241
  • Weisenbeck, D. R., Nelson, E. M., & McLeod, D. S. 2022. Herpetological inventory at Sweet’s Farm in Highland County, Virginia. Banisteria, 56, 118-136 - get paper here
  • Werler, John E. & James R. Dixon 2000. Texas Snakes. University of Texas Press, 544 pages
  • Werner, F. 1903. Neue Reptilien und Batrachier aus dem naturhistorischen Museum in Brüssel. Zool. Anz. 26: 246-253 - get paper here
  • Werning, Heiko 2012. Die Reptilien und Amphibien des Südwestens. Draco 13 (50): 18-60 - get paper here
  • Wilgers, D.J.; Horne, E.A.; Sandercock, B.K. & Volkmann, A.W. 2006. Effects of rangeland management on community dynamics of the herpetofauna of the Tall-grass Prairie [Flint Hills, Kansas/Oklahoma]. Herpetologica 62 (4): 378-388 - get paper here
  • Williams K L 1994. Lampropeltis triangulum (Lacepede), milk snake. Catalogue of American Amphibians and Reptiles 594: 1-10 - get paper here
  • Williams, K. L. 1978. Systematics and natural history of the American milk snake Lampropeltis triangulum. Milwaukee Publ. Mus. Contrib. Biol. Geol. No. 2, 258 pp. - get paper here
  • Williams, K. L. 1988. Systematics and natural history of the American milk snake Lampropeltis triangulum. 2nd ed. Milwaukee Publ. Mus. Contrib. Biol. Geol. - get paper here
  • Williams, Kenneth L. 1981. Literature: Systematics and natural history of the American milk snake, Lampropeltis triangulum. Litteratura Serpentium 1 (5): 206 - get paper here
  • Williams, M. & Foster, N. 2012. Geographic distribution: Lampropeltis triangulum (milksnake). Herpetological Review 43: 448-449 - get paper here
  • Williamson, Michael A. 1972. Lampropeltis triangulum celaenops. Herpetological Review 4 (5): 170 - get paper here
  • Woodbury, Angus M.;Hansen, Richard M. 1950. A snake den in Tintic Mountains, Utah. Herpetologica 6: 66-70 - get paper here
 
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