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Larutia kecil FUKUYAMA, HIKIDA, HOSSMAN & NISHIKAWA, 2019

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Higher TaxaScincidae, Sphenomorphinae (Sphenomorphini), Scincoidea, Sauria, Squamata (lizards)
Subspecies 
Common Names 
SynonymLarutia kecil FUKUYAMA, HIKIDA, HOSSMAN & NISHIKAWA 2019 
DistributionEast Malaysia (Borneo: Sarawak)

Type locality: west ridge of Gunung Penrissen, Kuching Division, Sarawak (N 1.1167, E 110.2147 (WGS84); 1,000 m asl.), East Malaysia, Borneo.  
Reproduction 
TypesHolotype. SRC 00624 (formerly, KUHE 60324), an adult male, collected by Ibuki Fukuyama and Kanto Nishikawa, on 30 March 2018. 
DiagnosisDiagnosis: Larutia kecil sp. nov. is a small species (SVL 84 mm) of Larutia with a relatively long head (HL/ SVL=0.13); body elongate; dorsal scales smooth; limbs poorly developed; lower eyelid bearing large, transparent scales, central scale largest; four supraoculars; frontoparietals paired; prefrontals small, widely separated; supra- nasals absent; postnasals absent; last supralabial horizontally divided or fused; first chinshields contacting second infralabial and each other medially; second chinshields separated from infralabials by an elongate scale and not in medial contact; 92 paravertebral scale rows; 22 longitudinal scale rows around midbody; caudal and body scales undifferentiated; dorsal head and body unicolor dark brown without yellow or pale bands or spots; rostral, first su- pralabial, nasal, frontonasal, mental, and first infralabial scales opaque.

Comparisons. Larutia kecil sp. nov. is clearly separated from all other species of Larutia by the number of subdigital lamellae beneath second toe (2 vs. 0 or 4 or 5; Fig. 6). The new species is further separated from all other species except of L. penangensis and L. seribuatensis in smaller adult body size (84 vs. 97–191); from all other spe- cies except L. miodactyla and L. sumatrensis, in the number of scales around midbody (22 vs. 18 or 23–30); from all other species except L. larutensis, L. miodactyla and L. nubisilvicola, in that the first two pairs of chinshields contact one infralabial on each side (vs. contacting two infralabials); from L. penangensis in that the second chinshield is separated from the infralabials by a scale (vs. contacting infralabials); from L. larutensis (in juveniles only), L. ser- ibuatensis, L. trifasciata, and L. nubisilvicola (in only one specimen) in the absence of nuchal bands (Fig. 3); from L. seribuatensis and some L. trifasciata in the absence of yellow spots on the head (Fig. 3); and from L. miodactyla, L. sumatrensis, and L. puehensis in having opaque scales on the snout (Fig. 6). Diagnostic characteristics are sum- marized across all species in Table 2 in Fukuyama et al. 2019). 
Comment 
EtymologyThe species name is from the Malay word “kecil”, meaning small, alluding to the small size of the new species. 
References
  • FUKUYAMA, IBUKI; TSUTOMU HIKIDA, MOHAMAD YAZID HOSSMAN, KANTO NISHIKAWA 2019. A new species of the genus Larutia (Squamata: Scincidae) from Gunung Penrissen, Sarawak, Borneo. Zootaxa 4661 (3): 522–532 - get paper here
  • Kwet, A. 2020. Liste der im Jahr 2019 neubeschriebenen Reptilien. Elaphe 2020 (3): 44-67
 
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