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Laudakia melanura BLYTH, 1854

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Higher TaxaAgamidae (Agaminae), Sauria, Iguania, Squamata (lizards)
SubspeciesLaudakia melanura lirata (BLANDORD 1874)
Laudakia melanura melanura BLYTH 1854
Laudakia melanura nasiri BAIG 1999 
Common NamesE: Black agama 
SynonymLaudakia (Placoderma) melanura BLYTH 1854: 738
Stellio melanurus — ANDERSON 1871: 189
Laudakia (Plocederma) melanura — ANDERSON 1871: 189 (in error)
Stellio liratus BLANDORD 1874 (fide SMITH 1935)
Agama melanura — BOULENGER 1885: 363
Agama lirata — BOULENGER 1885: 364
Agama melanura — HORA & CHOPRA 1923
Agama melanura — SMITH 1935: 218
Agama melanura — ANDERSON 1963: 475
Agama melanura HAAS & WERNER 1969
Agama melanura lirata — MERTENS 1969
Laudakia melanura — DAS 1996: 44
Laudakia melanura — BARTS & WILMS 2003
Laudakia lirata — KHAN 2004
Laudakia melanura lyrata — SINDACO & JEREMČENKO 2008
Laudakia melanura lirata — RAJABIZADEH & RASTEGAR-POUYANI 2009
Laudakia melanura lirata — HEIDARI & KAMI et al. 2009
Laudakia melanura lirata — HEIDARI & KAMI 2010
Laudakia lirata — KHAN et al. 2014
Laudakia melanura lirata — HABEEB 2018

Laudakia melanura melanura (BLYTH 1854)
Laudakia (Placoderma) melanura BLYTH 1854: 738
Laudakia melanura melanura — BAIG et al. 2012: 243

Laudakia melanura nasiri BAIG 1999
Laudakia melanura nasiri — SINDACO & JEREMČENKO 2008
Laudakia melanura nasiri — BAIG et al. 2012: 244 
DistributionIran, Pakistan, India (NW Punjab)

melanura: Pakistan (Salt Range in N Punjab, Waziristan in NW Frontier province, W Sind and Baluchistan provinces), SE Iran, SE Afghanistan. Type locality: Kashmir (Salt Range, Punjab, fide SMITH 1935)

nasiri: S Pakistan; Toba Kakar Range, Balochistan, Pakistan, elevation 2300-2400 m elevation. Type locality: “Tanishpa, Dist. Killa Saifullah, Toba Kakar Range, Balochistan, Pakistan; 31° 12’ N, 68° 28’ E, elevation 2320 m.” Only known from the type locality [BAIG et al. 2012].

lirata: Pakistan, Iran; Type locality: Gedrosia, Baluchistan (= Saman, Dasht Province, Baluchistan, fide SMITH 1935).  
TypesTypes: unlocated (ANDERSON 1999)
Holotype: ZSI 3410 [lirata]
Holotype: PMNH 735, female, collected by K. J. Baig 1997. [nasiri] 
DiagnosisDiagnosis (melanura). Size large; head and body dorsolaterally depressed; nostril on canthus rostralis; spinose scales on the sides of head greatly reduced; vertebral enlarged scales smooth or keeled; no patch of enlarged mucro- nate scales on flanks; tail long; tail segment inconsist- ent, starts with 2 – 3 whorls and gradually changes into 5, sometimes 6 in terminal part; callous glands present in males both at precloacal and abdominal positions [BAIG et al. 2012].

Diagnosis (nasiri): Medium size, head and body greatly de- pressed, vertebral scales enlarged, number of scales rows around midbody more than 150, no patch of spinose scales at the flanks, tail segmented, each segment with 2 to 4 rings. 
CommentSynonymy: mostly after BAIG et al. 2012.

Types: (for L. viridis var. woosnami) 
  • Ananjeva, N.B. & Tuniev 1994. Some aspects of historical biogeography of Asian rock agamids. Russ. J. Herpetol. 1 (1): 43 - get paper here
  • Anderson, J. 1871. On two Saurian genera Eurylepis and Plocederma Blyth, with a description of a new species of Mabouia Fitzinger. Proc. Asiat. Soc. Bengal 1871: 180-190 - get paper here
  • Anderson, S.C. 1963. Amphibians and Reptiles from Iran. Proc. Cal. Acad. Sci. Ser. 4, 31(16): 417-498 - get paper here
  • Anderson, Steven C 1999. The lizards of Iran. Contributions to Herpetology Volume 15, Society for the Study of Amphibians and Reptiles, Saint Louis, Missouri: i-vii, 1-442 [review in Copeia 2000 (4): 1144] - get paper here
  • Baig K J 1989. A new species of Agama (Sauria: Agamidae) from northern Pakistan. BULLETIN OF THE KITAKYUSHU MUSEUM OF NATURAL HISTORY (No. 9) 1989: 117-122
  • Baig, K. J. 1999. Description and ecology of a new subspecies of Black Rock Agama, Laudakia melanura (Sauria: Agamidae) from Balochistan, Pakistan. Russ. J. Herpetol. 6 (2): 81-86 - get paper here
  • Baig, Khalid Javed; Philipp Wagner, Natalia B. Ananjeva & Wolfgang Böhme 2012. A morphology-based taxonomic revision of Laudakia Gray, 1845 (Squamata: Agamidae). Vertebrate Zoology 62 (2): 213-260 - get paper here
  • Baig,K.J. & Böhme,W. 1995. Partition of the Stellio-group of Agama. 8th Ord. Gen. Meet. Soc. Europ. Herpet.: 36
  • Barts, M. & Wilms, T. 2003. Die Agamen der Welt. Draco 4 (14): 4-23 - get paper here
  • Blanford,W.T. 1874. Descriptions of new lizards from Persia and Baluchistàn. Ann. Mag. nat. Hist. (4) 13: 453-455 - get paper here
  • Blyth,E. 1854. Proceedings of the Society. Report of the Curator, Zoological Department. J. Asiat. Soc. Bengal 23 [1854]: 737-740 - get paper here
  • Boulenger, G.A. 1885. Catalogue of the lizards in the British Museum (Nat. Hist.) I. Geckonidae, Eublepharidae, Uroplatidae, Pygopodidae, Agamidae. London: 450 pp. - get paper here
  • Boulenger, George A. 1890. The Fauna of British India, Including Ceylon and Burma. Reptilia and Batrachia. Taylor & Francis, London, xviii, 541 pp. - get paper here
  • Haas G. and Werner, Y. L. 1969. Lizards and snakes from Southwestern Asia, collected by Henry Field. Bull. Mus. Comp. Zool. Harvard 138: 327-406. - get paper here
  • Habeeb, Israa Nadhim 2018. Check list lizards of Iraq. Mesopotamia Environmental Journal 4 (4): 25-33 - get paper here
  • Heidari, N.; H.G. Kami 2009. Lizards of the Gando Protected area in Sistan and Baluchestan Province, southeastern Iran. Iranian Journal of Animal Biosystematics 5 (2): 57-64 - get paper here
  • Heidari, Nastaran; Hamed Cheatsazan, Haji Gholi Kami, Soheila Shafiei 2010. Sexual dimorphism of the Black Rock Agama, Laudakia melanura lirata (Blanford, 1874) (Sauria: Agamidae). Zoology in the Middle East 49: 49-54 - get paper here
  • Hora, S.L.; Chopra, B. 1923. Reptilia and Batrachia of the Salt Range, Punjab. Records of the Indian Museum 4 (25): 369-376
  • Khan, M. S. 2014. Herpetology of Pakistan FIELD GUIDE SERIES: Families Agamidae, Chamaeleonidae. Privately published, 39 pp. - get paper here
  • Khan, M.S. 2003. CHECKLIST AND KEY TO THE LIZARDS OF PAKISTAN. Pakistan J. Zool. Suppl. Ser. (1): 1-25 - get paper here
  • Khan, Muhamad Sharif 2004. Annotated checklist of Amphibians and Reptiles of Pakistan. Asiatic Herpetological Research 10: 191-201 - get paper here
  • Rais, Muhammad; Ayesha Akram, Syeda Maria Ali, Muhammad Arslan Asadi, Misbah Jahangir, Muhammad Jawad Jilani, <br />and Maqsood Anwar 2015. Qualitative Analysis of Factors Influencing the Diversity and Spatial Distribution of Herpetofauna in Chakwal Tehsil (Chakwal District), Punjab, Pakistan. Herp. Cons. Biol. 10 (3) - get paper here
  • Rajabizadeh, M. & Rastegar-Pouyani, N. 2009. Two New Records of Reptiles (Reptilia: Squamata) from Southeastern Iran. Turkish Journal of Zoology 33: 103-104 - get paper here
  • RASTEGAR-POUYANI, NASRULLAH; HAJI GHOLI KAMI, MEHDI RAJABZADEH, SOHEILA SHAFIEI AND STEVEN CLEMENT ANDERSON 2008. Annotated Checklist of Amphibians and Reptiles of Iran. Iranian Journal of Animal Biosystematics 4 (1): 7-30
  • Schwier, H. 2008. Bemerkungen zum Geschlechtsdimorphismus bei der Schwarzagame (Laudakia melanura melanura BLYTH, 1854) anhand von zwei Abbildungen. Iguana Rundschreiben 20 (2): 23-24 [2007]
  • Sindaco, R. & Jeremcenko, V.K. 2008. The reptiles of the Western Palearctic. Edizioni Belvedere, Latina (Italy), 579 pp. - get paper here
  • ŠMÍD, JIŘÍ; JIŘÍ MORAVEC, PETR KODYM, LUKÁŠ KRATOCHVÍL, SEYYED SAEED HOSSEINIAN YOUSEFKHANI, ESKANDAR RASTEGAR-POUYANI & DANIEL FRYNTA 2014. Annotated checklist and distribution of the lizards of Iran. Zootaxa 3855 (1): 001–097 - get paper here
  • Smith, M.A. 1935. The fauna of British India, including Ceylon and Burma. Reptiles and Amphibia, Vol. II. Sauria. Taylor and Francis, London, 440 pp.
  • Venugopal, P.D. 2010. An updated and annotated list of Indian lizards (Reptilia: Sauria) based on a review of distribution records and checklists of Indian reptiles. Journal of Threatened Taxa 2 (3): 725-738. - get paper here
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