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Laudakia nuristanica (ANDERSON & LEVITON, 1969)

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Higher TaxaAgamidae (Agaminae), Sauria, Iguania, Squamata (lizards)
Subspecies 
Common NamesE: Leviton's Rock Agama 
SynonymAgama nuristanica ANDERSON & LEVITON 1969: 39
Stellio nuristanicus — ANANJEVA et al. 1990: 113
Laudakia nuristanica — ANANJEVA & TUNIEV 1994: 43
Laudakia nuristanica — SINDACO & JEREMČENKO 2008
Laudakia nuristanica — BAIG et al. 2012: 246
Laudakia nuristanica — JABLONSKI et al. 2021 
DistributionE Afghanistan (Nuristan, Kunar, Panjshir, Parwan, Takhar), NW Pakistan (Chitral Valley)

Type locality: Kamdesh, eastern Afghanistan, 1342 m elevation.  
Reproductionoviparous 
TypesHolotype: FMNH 161136, paratype: CAS 115939; other specimens: ZFMK, PMNH, UOCH (see list in Jablosnki et al. 2021); specimens reported from UF are actually Paralaudakia caucasia (P. Uetz & D. Jablonski, pers. comm.) 
DiagnosisDiagnosis: Head and body depressed; tail longer than head and body;
segmentation into whorls composed of four regular annuli barely distinct; about 45 scales around thickest part of tail; tympanum large, superficial; largest scales of back not arranged in regular rows and somewhat heterogeneous in size, a few about twice as large as largest ventral scales, strongly keeled, dorsolateral surfaces covered by minute granules (much smaller than found either in A. agrorensis or A. tuberculata flanks with scattered enlarged scales, not grouped in patches, small scales granular; limbs above covered with strongly hetero geneous scales, numerous greatly enlarged scales among much smaller ones, the smaller almost granular; large scales neither grouped in patches nor imbricate, but strongly spinose (Anderson & Leviton 1969: 40).

Diagnosis. This form is very close to L. tuberculata and may be distinguished by having a higher number of scales around the body and tail, indistinct tail segment, and a unique type of scattered, enlarged conical, spinose scales over the limbs and body [BAIG et al. 2012]. 
CommentAbundance: rare; Only known from 17 specimens in five museum collections. 
EtymologyNamed after the type locality in Nuristan Province. 
References
  • Ananjeva, N.B. & Tuniev 1994. Some aspects of historical biogeography of Asian rock agamids. Russ. J. Herpetol. 1 (1): 43 - get paper here
  • Ananjeva, N.B.; Peters,G.; Macey & Papenfuss, T. 1990. Stellio sacra (Smith 1935) - a distinct species of Asiatic rock agamid from Tibet. Asiatic Herpetological Research 3: 104-115 - get paper here
  • Anderson, S. C. & A. E. Leviton 1969. Amphibians and reptiles collected by the Street expedition to Afghanistan, 1965. Proc. Cal. Acad. Sci. (4) 37: 25-56 - get paper here
  • Baig, K.J. 1988. New record of Agama nuristanica (Sauria: Agamidae) from Pakistan. Biologia, Islamabad, 34: 199-200
  • Baig, Khalid Javed; Philipp Wagner, Natalia B. Ananjeva & Wolfgang Böhme 2012. A morphology-based taxonomic revision of Laudakia Gray, 1845 (Squamata: Agamidae). Vertebrate Zoology 62 (2): 213-260 - get paper here
  • Baig,K.J. & Böhme,W. 1995. Partition of the Stellio-group of Agama. 8th Ord. Gen. Meet. Soc. Europ. Herpet.: 36
  • Jablonski, D., Basit, A., Farooqi, J., & Masroor, R. 2021. Distribution and morphological characters of Laudakia nuristanica (Anderson & Leviton, 1969) from Afghanistan. Zoology in the Middle East 67(3): 240–246. - get paper here
  • Khan, M. S. 2014. Herpetology of Pakistan FIELD GUIDE SERIES: Families Agamidae, Chamaeleonidae. Privately published, 39 pp. - get paper here
  • Khan, M.S. 2003. CHECKLIST AND KEY TO THE LIZARDS OF PAKISTAN. Pakistan J. Zool. Suppl. Ser. (1): 1-25 - get paper here
  • Sindaco, R. & Jeremcenko, V.K. 2008. The reptiles of the Western Palearctic. Edizioni Belvedere, Latina (Italy), 579 pp. - get paper here
 
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