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Leiocephalus macropus (COPE, 1863)

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Higher TaxaLeiocephalidae, Iguania, Sauria, Squamata (lizards)
SubspeciesLeiocephalus macropus macropus COPE 1863
Leiocephalus macropus aegialus SCHWARTZ & GARRIDO 1967
Leiocephalus macropus asbolomus SCHWARTZ & GARRIDO 1967
Leiocephalus macropus felinoi GARRIDO 1979
Leiocephalus macropus hoplites ZUG 1959
Leiocephalus macropus hyacinthurus ZUG 1959
Leiocephalus macropus immaculatus HARDY 1958
Leiocephalus macropus koopmani ZUG 1959
Leiocephalus macropus lenticulatus GARRIDO 1973
Leiocephalus macropus phylax SCHWARTZ & GARRIDO 1967
Leiocephalus macropus torrei GARRIDO 1979 
Common NamesE: Cuban Side-blotched Curlytail, Monte Verde Curlytail Lizard 
SynonymLiocephalus (sic) macropus COPE 1863:184
Liocephalus macropus — BOULENGER 1885: 163
Leiocephalus macropus — STEJNEGER 1917
Leiocephalus macropus — SCHWARTZ & HENDERSON 1991: 435
Leiocephalus macropus — RODRÍGUEZ SCHETTINO 1999: 135

Leiocephalus macropus aegialus SCHWARTZ & GARRIDO 1967
Leiocephalus macropus aegialus — PREGILL 1992: 47
Leiocephalus macropus aegialus — HEDGES et al. 2019

Leiocephalus macropus asbolomus SCHWARTZ & GARRIDO 1967
Leiocephalus macropus asbolomus — PREGILL 1992: 47
Leiocephalus macropus asbolomus — HEDGES et al. 2019

Leiocephalus macropus felinoi GARRIDO 1979
Leiocephalus macropus felinoi — PREGILL 1992: 47
Leiocephalus macropus felinoi — HEDGES et al. 2019

Leiocephalus macropus hoplites ZUG 1959
Leiocephalus macropus hoplites — PREGILL 1992: 47
Leiocephalus macropus hoplites — HEDGES et al. 2019

Leiocephalus macropus hyacinthurus ZUG 1959
Leiocephalus macropus hyacinthurus — PREGILL 1992: 47
Leiocephalus macropus hyacinthurus — HEDGES et al. 2019

Leiocephalus macropus immaculatus HARDY 1958
Leiocephalus macropus immaculatus — PREGILL 1992: 47
Leiocephalus macropus inmaculatus [sic] — RODRÍGUEZ SCHETTINO 1999: 137
Leiocephalus macropus immaculatus — HEDGES et al. 2019

Leiocephalus macropus koopmani ZUG 1959
Leiocephalus macropus koopmani — PREGILL 1992: 47
Leiocephalus macropus koopmani — HEDGES et al. 2019

Leiocephalus macropus lenticulatus GARRIDO 1973: 10
Leiocephalus macropus lenticulatus — PREGILL 1992: 47
Leiocephalus macropus lenticulatus — HEDGES et al. 2019

Leiocephalus macropus phylax SCHWARTZ & GARRIDO 1967
Leiocephalus macropus phylax — PREGILL 1992: 47
Leiocephalus macropus phylax — HEDGES et al. 2019

Leiocephalus macropus torrei GARRIDO 1979
Leiocephalus macropus torrei — PREGILL 1992: 47
Leiocephalus macropus torrei — HEDGES et al. 2019 
DistributionCuba (from Pinar del Rio east, but primarily E Cuba)

macropus: SE coast from Bahia de Santiago east at least to Sierra del Purial north of Cajobabo; presumably inland to type-locaiity in Sierra del Guaso; Isla de la Juventud? Type locality: Eastern Cuba; restricted by Stejneger, 1917, to Monte Verde, Guantánamo Province, Cuba.

aegialus: northern coastal Camagüey Province.

asbolomus: northeastern coast of Guantânamo Province, west to Holguin Province.

felinoi: north coastal Matanzas Province.

hoplites: northern Ciego de Âvila Province.

hyacinthurus: Sierra de Escambray.

immaculatus: southeastern coast from Rio Magdalena east to Bahia de Santiago; Cayo Damas.

koopmani: Peninsula de Guanahacabibes.

lenticulatus: coastal hills west of Gibara, Holguin Province. Type locality: “arenal Los Cocos, a seis km de Gibara”

phylax: southwestern Granma Province, from Verre6n east to Rio Puercos and Punta Hicacos.

torrei: along Rio Paredones, Matanzas Province.

INCERTAE SEDIS: Material from the following localities has not been assigned to subspecies: Niquero, Cayo Saetia, Pinares de Mayari, Sierra de Nipe, Cupeyai, all in eastern Cuba; Miranda, Santiago de Cuba Province; Rangel, Pinar del Rio Province; Punta Hicacos, Matanzas Province.  
Reproductionoviparous 
TypesLectotype: USNM 25819, selected by Hardy, 1958; Syntypes: MCZ 10930; USNM 12254; USNM 25819-23, USNM 25825-26.
Holotype: AMNH 83255 [aegialus]
Holotype: CZACC = IZ 568 [asbolomus]
Holotype: CZACC = IZ 4751 [felinoi]
Holotype: AMNH 78020 [hoplites]
Holotype: AMNH 78015 [hyacinthurus]
Holotype: USNM 138412 [immaculatus]
Holotype: MCZ 55541 [koopmani]
Holotype: CZACC = IZ 2782 [lenticulatus]
Holotype: CZACC = IZ 556 [phylax]
Holotype: CZACC = IZ 3725 [torrei] 
DiagnosisAdditional details, e.g. a detailed description or comparisons (3575 characters), are available for collaborators and contributors. Please contact us for details. 
CommentSee Schwartz and Garrido, 1967, for comments on restricted type-locality, status of syntypes, and selection of lectotype.

Distribution: see map of subspecies in RODRÍGUEZ SCHETTINO 1999: 136. 
References
  • Alfonso, Yasel U.; Pierre Charruau, Gabriel Fajardo and Alberto R. Estrada. 2012. Interspecific communal oviposition and reproduction of three lizard species in Southeastern Cuba. Herpetology Notes 5: 73-77.
  • Bakkegard, Kristin A.; David A. Johnson, and Daniel G. Mulcahy 2020. A New Locality, Naval Station Guantanamo Bay, for the Rare Lizard Leiocephalus onaneyi (Guantánamo Striped Curlytail) and Notes on its Natural History. CARIBBEAN NATURALIST (79): 1–22
  • Boulenger, G.A. 1885. Catalogue of the lizards in the British Museum (Natural History). Vol. 2, Second edition. London, xiii+497 pp. - get paper here
  • Cope, E.D. 1863. Contributions to Neotropical saurology. Proc. Acad. Nat. Sci. Philadelphia 14 [1862]: 176-188. - get paper here
  • Garrido, O. H. 1973. Nueva especie de Leiocephalus (Lacertilia, Iguanidae) para Cuba. Poeyana 116:1-19.
  • Garrido, O.H. 1973. Nuevas subespecies de reptiles para Cuba. Torreia, n.s. 30:1-31.
  • Garrido, O.H. 1979. Nuevas subespecies de Leiocephalus macropus Cope (Lacertilia: Iguanidae) para Cuba. Poeyana (188):1-16.
  • Hardy, J.D., Jr. 1958. A geographic variant gradient in the Cuban lizard Leiocephalus macropus Cope. Herpetologica 13 (3): 275-76 - get paper here
  • Hedges SB, Powell R, Henderson RW, Hanson S, and Murphy JC 2019. Definition of the Caribbean Islands biogeographic region, with checklist and recommendations for standardized common names of amphibians and reptiles. Caribbean Herpetology 67: 1–53
  • Pregill,G.K. 1992. Systematics of the West Indian Lizard Genus Leiocephalus (Squamata: Iguania: Tropiduridae). Misc. Publ. Univ. Kans. Mus. Nat. Hist. 84: 1-69 - get paper here
  • Rodríguez Schettino, Lourdes (ed.) 1999. The iguanid lizards of Cuba. University Press of Florida, 428 pp.
  • Rodríguez Schettino, Lourdes, Carlos A. Mancina & Vilma Rivalta González 2013. REPTILES OF CUBA: CHECKLIST AND GEOGRAPHIC DISTRIBUTIONS. Smithsonian Herp. Inf. Serv. (144): 1-96 - get paper here
  • Schwartz, A. & Henderson, R.W. 1991. Amphibians and Reptiles of the West Indies. University of Florida Press, Gainesville, 720 pp.
  • Schwartz, A. and Garrido, O.H. 1967. A review of the Cuban lizard Leiocephalus macropus Cope. Reading Public Museum and Art Gallery Sci. Publ. (14): 1-41.
  • Zug, G.R. 1959. Three new subspecies of the lizard Leiocephalus macropus Cope from Cuba. Proc. Biol. Soc. Washington72:139-50. - get paper here
 
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