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Leiolepis triploida PETERS, 1971

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Higher TaxaAgamidae (Leiolepididae or Leiolepidinae), Sauria, Iguania, Squamata (lizards)
Subspecies 
Common NamesE: Thai Butterfly Lizard, Malaysian Butterfly Lizard
G: Malayische Schmetterlingsagame 
SynonymLeiolepis triploida PETERS 1971: 123
Leiolepis triploida — MANTHEY & GROSSMANN 1997: 286
Leiolepis triploida — CHAN-ARD et al. 1999: 134
Leiolepis triploida — GRISMER 2011
Leiolepis triploida — GRISMER & QUAH 2019 
DistributionThailand, W Malaysia

Type locality: Malayisch-thailändisches Grenzgebiet auf der Malayischen Halbinsel.  
Reproductiontriploid, parthenogenetic species. Seems to have a complex parental history with multiple hybridizations between L. guttata (M) × L. reevesii (GRISMER et al. 2014). 
TypesHolotype: MCZ 104381; 28 Paratypes in ZMB, MCZ, ZFMK (ZFMK 30698 ehem. MCZ 101923). 
DiagnosisDiagnosis (original): “Eine durch Triploidie (3n = 54) und ameiotische Parthenogenese von ihrer Ursprungsform (Leiolepis belliana) isolierte Agamospecies von beschränkter Fertilität. In der Rückenzeichnung L. belliana ähnlich: es sind 3 helle kontinuierliche Längsstreifen vorhanden (der mittlere ist stets blasser alsdie äußeren), doch die fahlen Ozellenflecken zwischen den Streifen stehen eng beieinander und ihre dunklen Bänder bilden ein polygonales Maschenwerk. Keine Fabkontrastzeichnung an den Hals- und Rumpfseiten; stattdessen nur unscharfe Querstreifen oder -tüpfel von schwefel bis orangegelber Tönung. In der Pholidose vor allem durch die relativ kleinen Buchschilder von L. belliana abweichend (17-19,4 - 22 gegenüber 10-15,2 - 18 pro Zählstrecke bei L. belliana). Sehr großwüchsig und auffallend langschwänzig.” 
Comment 
EtymologyNamed after its triploid karyotype. 
References
  • Chan-ard,T.; Grossmann,W.; Gumprecht,A. & Schulz,K. D. 1999. Amphibians and reptiles of peninsular Malaysia and Thailand - an illustrated checklist [bilingual English and German]. Bushmaster Publications, Würselen, Gemany, 240 pp. [book review in Russ. J Herp. 7: 87] - get paper here
  • Darevsky I S; Kupriyanova L A 1993. Two new all-female lizard species of the genus Leiolepis Cuvier, 1829 from Thailand and Vietnam (Squamata: Sauria: Uromastycinae). Herpetozoa 6 (1-2): 3-20 - get paper here
  • Grismer, J.L. & Grismer, L.L. 2010. Who’s your mommy? Identifying maternal ancestors of asexual species of Leiolepis Cuvier, 1829 and the description of a new endemic species of asexual Leiolepis Cuvier, 1829 from Southern Vietnam. Zootaxa 2433: 47–61 - get paper here
  • Grismer, Jesse L.; Aaron M. Bauer, L. Lee Grismer, Kumthorn Thirakhupt, Anchelee Aowphol, Jamie R. Oaks, Perry L. Wood Jr, Chan Kin Onn, Neang Thy, Micheal Cota and Todd Jackman 2014. Multiple origins of parthenogenesis, and a revised species phylogeny for the Southeast Asian butterfly lizards, Leiolepis. Biological Journal of the Linnean Society 113 (4): 1080–1093 - get paper here
  • GRISMER, L. LEE; & EVAN S. H. QUAH 2019. An updated and annotated checklist of the lizards of Peninsular Malaysia, Singapore, and their adjacent archipelagos. Zootaxa 4545 (2): 230–248 - get paper here
  • Grismer, L.L. 2011. Lizards of Peninsular Malaysia, Singapore and their adjacent archipelagos. Edition Chimaira, Frankfurt, 728 pp. [review in Herp. Rev. 43: 155] - get paper here
  • Manthey U 2010. Agamid Lizards of Southern Asia. Draconinae 2 -Leiolepidinae. Edition Chimaira, Terralog 7b, Frankfurt, 168 pp.
  • Manthey, U. & Grossmann, W. 1997. Amphibien & Reptilien Südostasiens. Natur und Tier Verlag (Münster), 512 pp. - get paper here
  • ONN, CHAN KIN; L. LEE. GRISMER, DIONYSIUS S. SHARMA, DAICUS BELABUT, and NORHAYATI AHMA 2009. New herpetofaunal records for Perlis State Park and adjacent areas. Malayan Nature Journal 61 (4): 255 - 262
  • Peters, G. 1971. Die intragenerischen Gruppen und die Phylogenese der Schmetterlingsagamen (Agamidae: Leiolepis). Zool. Jahrb. (Syst.) 98: 11-130
  • Talib, A., Hambali, K., Ngadi, E., Haiqal, M. S., Amir, A., & Yin, S. A. 2020. A preliminary survey of herpetofauna at Delta Tumpat Mangrove Forest, Kelantan, Malaysia. Journal of Wildlife and Parks - get paper here
  • Zakaria, N., Allahudin, M. I. H., Ma'ad, S. N. S., Sulaiman, A. A., Abdullah, N. A., Zamri, M. I. M., ... & Deraman, M. Y. 2022. Diversity amphibians and reptiles at Sungai Kerteh mangrove forest, Terengganu, Malaysia. Biodiversitas Journal of Biological Diversity, 23(11)
 
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