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Limaformosa guirali (MOCQUARD, 1887)

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Higher TaxaLamprophiidae, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies 
Common NamesE: Mocquard's File Snake 
SynonymHeterolepis guirali MOCQUARD 1887: 23
Heterolepis bicarinatus SAUVAGE 1884: 145 (non DUMÉRIL & BIBRON)
Simocephalus guirali – BOULENGER 1893: 346
Simocephalus baumanni STERNFELD 1908: 93 (fide HALLERMANN 2007)
Mehelya guirali – LÖNNBERG 1910
Mehelya (S.) baumanni – LÖNNBERG 1910
Simocephalus guirali — WITTE 1933
Mehelya guirali — LOVERID1939: 136
Simocephalus guirali — ROMER 1953
Mehelya guirali — TRAPE & ROUX-ESTÈVE 1995: 39
Mehelya guirali — BURGER et al. 2004
Gonionotophis guirali — KELLY et al. 2011
Mehelya guirali — SEGNIAGBETO et al. 2012: 344
Mehelya guirali — HUGHES 2013
Mehelya guirali — WALLACH et al. 2014: 425
Mehelya guirali — TRAPE & BALDÉ 2014
Limaformosa guirali — BROADLEY et al. 2018
Gonionotophis guirali — CHIPPAUX & JACKSON 2019: 214
Gonionotophis guirali — SENTER & CHIPPAUX 2022 
DistributionLiberia, Sierra Leone, Ivory Coast, Ghana, Togo, Benin ?, Guinea (Conakry), Nigeria, Cameroon, Central African Republic, N Zaire, Gabon, Congo (Brazzaville) ?, Equatorial Guinea

Type locality: Niger River, Nigeria.  
Reproductionoviparous 
TypesSyntypes: MNHN-RA 1885.0580 (MNHN-RA 1885.0580A is the head skull of the same specimen) and MNHN-RA 1883.0591, one being an 1130 mm (SVL) and 1060 mm specimen (Lanchier and L. Guiral)
Holotype: ZMB 2197615 from Togo [Simocephalus baumanni] 
Diagnosis 
CommentDistribution: Not listed by BROADLEY & POYNTON 1998 for Democratic Republic of the Congo (Zaire). Not reported from Benin according to ULLENBRUCH et al. 2010 but confirmed by HUGHES 2013. For a map with localities in Equatorial Guinea see SÁNCHEZ-VIALAS et al. 2022. 
EtymologyNamed after Leon Guiral (1858-1885), former French naval quartermaster and a member of de Brazza's expeditions in the French Congo. 
References
  • Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here
  • Böhme, Wolfgang, Mark-Oliver Rödel, Christian Brede & Philipp Wagner 2011. The reptiles (Testudines, Squamata, Crocodylia) of the forested southeast of the Republic Guinea (Guinée forestière), with a country-wide checklist. Bonn zoological Bulletin 60 (1): 35-61 - get paper here
  • Boulenger, G.A. 1893. Catalogue of the snakes in the British Museum (Nat. Hist.) I. London (Taylor & Francis), 448 pp. - get paper here
  • Broadley, Donald G.; Krystal A. Tolley, Werner Conradie, Sarah Wishart, Jean-François Trape, Marius Burger, Chifundera Kusamba, Ange-Ghislain Zassi-Boulou & Eli Greenbaum 2018. A phylogeny and genus-level revision of the African file snakes Gonionotophis Boulenger (Squamata: Lamprophiidae). African Journal of Herpetology, DOI: 10.1080/21564574.2018.1423578 - get paper here
  • Burger, M.; Branch, W.R. & Channing, A. 2004. Amphibians and Reptiles of Monts Doudou, Gabon: Species Turnover Along an Elevational Gradient. California Academy of Sciences Memoir 28: 145–186
  • Chippaux, Jean-Philippe & Kate Jackson 2019. Snakes of Central and Western Africa. Johns Hopkins University Press, 448 pp. [detaileld review in HR 51 (1): 161] - get paper here
  • Chirio, L. & Lebreton, M. 2007. Atlas des reptiles du Cameroun. MNHN, IRD, Paris 688 pp.
  • Hallermann, J. 2007. The status of problematic snake types of the Museum für Naturkunde der Humboldt-Universität zu Berlin (Reptilia, Serpentes). Mitt. Mus. Nat.kd. Berl., Zool. Reihe 83 (2): 160–165 - get paper here
  • Hughes, B. 2013. Snakes of Bénin, West Africa. Bull. Soc. Herp. France 144: 101-159
  • Kelly, Christopher M.R.; William R. Branch, Donald G. Broadley, Nigel P. Barker, Martin H. Villet 2011. Molecular systematics of the African snake family Lamprophiidae Fitzinger, 1843 (Serpentes: Elapoidea), with particular focus on the genera Lamprophis Fitzinger 1843 and Mehelya Csiki 1903. Mol Phylogenet Evol. 58(3): 415-426 - get paper here
  • Lönnberg, E. 1910. Two new snakes from Lower Congo. Arkiv för Zoologi 7 (8): 1-8 - get paper here
  • Loveridge, A. 1938. On a collection of reptiles and amphibians from Liberia. Proc. New England Zool. Club 17: 49-74
  • Loveridge, A. 1939. Revision of the african snakes of the genera Mehelya and Gonionotophis. Bull. Mus. Comp. Zool. Harvard 86: 131-162 - get paper here
  • Menzies, J.I. 1966. The snakes of Sierra Leone. Copeia 1966 (2): 169-179. - get paper here
  • Mocquard, F. 1887. Du genre Heterolepis et des espèces qui le composent, dont trois nouvelles. Bull. Soc. Philom. Paris (7) 11: 1-34 - get paper here
  • Pauwels, O.S.G. & Vande Weghe, J.P. 2008. Les reptiles du Gabon. Smithsonian Institution, Washington: 272 pp. - get paper here
  • PAUWELS, OLIVIER; LAURENT CHIRIO & WOUTER DEKONINCK. 2022. Diet records for snakes from Guinea, West Africa. Bulletin of the Chicago Herpetological Society 57(6): 117–123. - get paper here
  • Portik DM, Jongsma GFM, Kouete MT, Scheinberg LA, Freiermuth B, Tapondjou WP, Blackburn DC. 2016. A survey of amphibians and reptiles in the foothills of Mount Kupe, Cameroon. Amphibian & Reptile Conservation 10(2) [Special Section]: 37–67 (e131)
  • Romer, J.D. 1953. Reptiles and Amphibians Collected in the Port Harcourt Area of Nigeria. Copeia 1953 (2): 121-123 - get paper here
  • SÁNCHEZ-VIALAS, A., CALVO-REVUELTA, M. & DE LA RIVA, I. 2022. Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197 - get paper here
  • Sauvage, H.-E. 1884. Sur quelques Reptiles de la collection du Muséum d’Histoire Naturelle. Bull. Soc. Philom. Paris (7) 8: 142-146 - get paper here
  • Segniagbeto GLazcano. H., Trape J. F., David P., Ohler A., Dubois A. & Glitho I. A. 2011. The snake fauna of Togo: systematics, distribution and biogeography, with remarks on selected taxonomic problems. Zoosystema 33 (3): 325-360. DOI: 10.5252/z2011n3a4 - get paper here
  • Senter, P. J., & Chippaux, J. P. 2022. Biogeography of snakes in Liberia: Review and synthesis of current knowledge. Ghana Journal of Science, 63(1), 29-62 - get paper here
  • Sinsin B, Bergmans W(eds) 1999. Rongeurs, ophidiens et relations avec l’environnement agricole du Bénin. Cotonou (Éd. Flamboyant), 199 pp.
  • Sternfeld, R. 1908. Neue und ungenügend bekannte afrikanische Schlangen. S. Ber. Ges. naturforsch. Freunde Berlin, 4: 92-95 - get paper here
  • Trape, J.F. & R. ROUX-ESTÈVE 1995. Les serpents du Congo: liste commentée et clé de détermination. Journal of African Zoology 109 (1): 31-50
  • TRAPE, JEAN-FRANÇOIS & CELLOU BALDÉ 2014. A checklist of the snake fauna of Guinea, with taxonomic changes in the genera Philothamnus and Dipsadoboa (Colubridae) and a comparison with the snake fauna of some other West African countries. Zootaxa 3900 (3): 301–338 - get paper here
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp.
  • Witte, G. F. de 1933. Reptiles récoltés au Conge Belge par le Dr. H. Schouteden et par M. G.-F. Witte. Ann. Mus. Conge belge Zool. Ser. 1 Tome III: 53-100.
 
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