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Liolaemus puritamensis NÚÑEZ & FOX, 1989

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Higher TaxaLiolaemidae, Iguania, Sauria, Squamata (lizards)
Subspecies 
Common Names 
SynonymLiolaemus puritamensis NÚÑEZ & FOX 1989
Liolaemus (Eulaemus) puritamensis — RAMIREZ LEYTON & PINCHEIRA-DONOSO 2005: 141
Liolaemus (Eulaemus) filiorum RAMIREZ LEYTON & PINCHEIRA-DONOSO 2005
Liolaemus (Eulaemus) puritamensis — PINCHEIRA-DONOSO & NÚÑEZ 2005: 118
Liolaemus puritamensis — ABDALA et al. 2008
Liolaemus filiorum — LIVIGNI 2013: 362
Liolaemus puritamensis — TRONCOSO-PALACIOS 2015 
DistributionChile, Argentina (Jujuy, Puna region, 3500–4900 m elevation)

Type locality: “on the road to El Tatio springs (42 km northeast of San Pedro de Atacama)".

filiorum: Chile (El Loa: Antofagasta); Type locality: Cerro Las Papas, sur de Lequena, Provincia de El Loa, Segunda Regíon de Antofagasta, Chile  
ReproductionVivparous. 
TypesHolotype: MNHNC 1880 (Mus. Nac. Hist. Nat. Chile, Santiago)
Holotype: MNHN 3829 (also a paratype of L. puritamensis, fide Troncoso-Palacios 2015) [filiorum] 
DiagnosisDiagnosis: “This species differs from all known Chilean forms in having: a) coarse dorsal scales with a semirounded edge and a light keel, a character shared in part with the species of the genus Velosaura(Nuñez and Yañez, 1983-1984), but in L. puritamensis the scales are narrower; b) a dark venter spotted with white scales, shared with patagonian Liolaemus, but the latter do not exhibit the same spotted pattern; c) neck wider than head; d) dorsal scales, even though triangular with a free point, interspersed with various minute scales; and e) sides of body with smaller, rounder, more conical, protuberant scales than those of dorsum, also with interspersed minute scales.” (NÚÑEZ & FOX 1989) 
CommentSynonymy: NUNEZ 1992 synonymized L. puritamensis with L. dorbignyi, but resurrected by  PINCHEIRA-DONOSO & NÚÑEZ 2005, a decision accepted by LOBO et al. 2010. Troncoso-Palacio (2014) concludes that Liolaemus puritamensis Nuñez and Fox, 1989 and L. filiorum Pincheira-Donoso & Ramirez, 2005 are conspecific. However, this was challenged by Pincheira-Donoso 2014 and rebutted again by Troncoso-Palacios 2015.

Common name (filiorum): S: Lagartija de los hijos; E: Son’s lizard. 
EtymologyNamed after the nearby springs of Baños de Puritama.

L. filiorum was named after Ramirez-Leyton’s sons, Marcelo and Alvaro. Latin “filius” = son. 
References
  • Abdala CS, Aguilar-Kirigin AJ, Semhan RV, Bulacios Arroyo AL, Valdes J, Paz MM, et al. 2019. Description and phylogeny of a new species of Liolaemus (Iguania: Liolaemidae) endemic to the south of the Plurinational State of Bolivia. PLoS ONE 14(12): e0225815 - get paper here
  • Abdala, C.S.; S. Quinteros & R.E. Espinoza 2008. Two new species of Liolaemus (iguania: Liolaemidae) from the Puna of Northwestern Argentina. Herpetologica 64 (4): 458–471 - get paper here
  • AVILA, LUCIANO JAVIER; LORENA ELIZABETH MARTINEZ & MARIANA MORANDO 2013. Checklist of lizards and amphisbaenians of Argentina: an update. Zootaxa 3616 (3): 201–238 - get paper here
  • Demangel, Diego 2016. Reptiles en Chile. Fauna Nativa Ediciones, Santiago, 619 pp - get paper here
  • Demangel, Diego; Cristian Sepúlveda, Manuel Jara, Daniel Pincheira-Donoso and Herman Núñez 2015. Liolaemus omorfi, A NEW LIZARD SPECIES FROM THE ANDES OF NORTHERN CHILE (SAURIA, LIOLAEMIDAE). Boletín del Museo Nacional de Historia Natural, Chile, 64: 139-155 - get paper here
  • Langstroth, R.P. 2005. Adiciones probables y confirmadas para la saurofauna boliviana. Kempffiana 1 (1): 101-128
  • LiVigni, F. (ed.) 2013. A Life for Reptiles and Amphibians, Volume 1. Chimaira, Frankfurt, 495 pp. - get paper here
  • Lobo, F.; Espinoza, R.E. and S. Quinteros 2010. A critical review and systematic discussion of recent classification proposals for liolaemid lizards. Zootaxa 2549: 1–30 - get paper here
  • Nunez H; Fox S F 1989. Liolaemus puritamensis, a new species of iguanid lizard previously confused with Liolaemus multiformis (Squamata: Iguanidae). Copeia 1989 (2): 456-460 - get paper here
  • Pincheira-Donoso, Daniel 2014. No evidence for conspecificity between two high Andes Liolaemus lizards (Squamata: Liolaemidae). Acta Herpetologica 9 (2): 249-252 - get paper here
  • PINCHEIRA-DONOSO, Daniel & HERMAN NÚÑEZ 2005. The Chilean species of the genus Liolaemus Wiegmann, 1834 (Iguania, Tropiduridae, Liolaeminae). Taxonomy, systematics and evolution [in Spanish] Mus Nac Hist Nat Chile Publ Ocas 59: 1-486 - get paper here
  • QUINTEROS, ANDRÉS SEBASTIÁN & CRISTIAN SIMÓN ABDALA 2011. A new species of Liolaemus of the Liolaemus montanus section (Iguania: Liolaemidae) from Northwestern Argentina. Zootaxa 2789: 35–48 - get paper here
  • Ramírez Leyton, G. & Pincheira Donoso, D. 2005. Fauna del Altiplano y Desierto de Atacama. Phrynosaura Ediciones, 395 pp.
  • Troncoso-Palacio, J. 2014. Revisión del estatus taxonómico de Liolaemus filiorum Pincheira-Donoso y Ramírez, 2005 (Iguania: Liolaemidae). Cuad. Herpetol. 28: 1-7 - get paper here
  • Troncoso-Palacios, J. 2015. On the taxonomic status of Liolaemus filiorum Pincheira-Donoso & Ramírez, 2005 (Iguania: Liolaemidae): A response to Pincheira-Donoso. Acta Herpetologica 10 (2): 125-127; DOI: 10.13128/Acta_Herpetol-15394 - get paper here
 
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