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Liolaemus velosoi ORTIZ, 1987

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Higher TaxaLiolaemidae, Iguania, Sauria, Squamata (lizards)
Subspecies 
Common Names 
SynonymLiolaemus velosoi ORTIZ 1987
Liolaemus josephorum NÜÑEZ, SCHULTE & GARÍN 2001
Liolaemus (Liolaemus) velosoi — PINCHEIRA-DONOSO & NÚÑEZ 2005: 429
Liolaemus josephorum — LOBO et al. 2010
Liolaemus josephorum — TRONCOSO-PALACIOS & FERRI-YÁNEZ 2012
Liolaemus nigrocoeruleus MARAMBIO-ALFARO & TRONCOSO-PALACIOS 2014
Liolaemus velosoi — ESQUERRÉ et al. 2022
Liolaemus nigrocoeruleus — ÁVILA2 & MARAMBIO-ALFARO 2023 
DistributionChile

Type locality: “Desvio Cerro Imán près de Copiapó (27° 20’ S-70° 30’)".

josephorum: N Chile (Atacama region); Type locality: Diego de Almagro (26° 23’ S. 70° 02’ W, 750 msnm).

nigrocoeruleus: Chile (coastal Atacama desert); Type locality: siete kilómetros al NE de Barranquilla (27°27’35’’S – 70°50’43’’W, en una loma sin nombre, a 200 m sobre el nivel del mar. Provincia de Copiapó, Región de Atacama, Chile.  
Reproductionoviparous 
TypesHolotype: MZUC 10897 (Mus. Zool. Univ. Concepción)
Holotype: MNHN 2532 [josephorum]
Holotype: SSUC Re 0527 (Fig. 1), adult male. Colectado por J. Troncoso-Palacios y Y. Marambio-Alfaro en mayo de 2012.
Paratipos: SSUC Re 0528 a 0532, 0552 a 0554. Dos machos, tres juveniles y tres hembras. Colectados a siete kilómetros al NE de Barranquilla (27°27’35’’S – 70°50’43’’W, Fig. 2), en una loma sin nombre, a 200 m sobre el nivel del mar. Provincia de Copiapó, Región de Atacama, Chile. Colectados por J. Troncoso-Palacios y Y. Marambio-Alfaro en mayo de 2012. SSUC Re 0533 y 0534. Un macho y un juvenil. Colectados a nueve kiló- metros al NE de Barranquilla (27°27’00’’S – 70°49’41’’W), en una loma sin nombre, a 180 m sobre el nivel del mar. Provincia de Copiapó, Región de Atacama, Chile. Colectados por J. Troncoso-Palacios y Y. Marambio-Alfaro en mayo de 2012 [nigrocoeruleus] 
DiagnosisAdditional details, e.g. a detailed description or comparisons (2336 characters), are available for collaborators and contributors. Please contact us for details. 
CommentSynonymy: PINCHEIRA-DONOSO et al. 2008 synonymized josephorum with velosoi, a decision that is not supported by LOBO et al. 2010 (although the latter do not list the species in their list!), but confirmed by TRONCOSO-PALACIOS & FERRI-YÁNEZ 2012. Synonymized with platei by Demangel 2016. ESQUERRÉ et al. 2022 synonymized Liolaemus nigrocoeruleus with L. velosoi. 
EtymologyNamed after Alberto Veloso for his contributions to the knowledge of Chiliean frogs.

Liolaemus nigrocoeruleus was named after the black (”nigro”) and blue (Latin ”caeruleus”) coloration of males. 
References
  • Ávila, J. M., & Marambio-Alfaro, Y. 2023. Reptiles of the Atacama Region, northern Chile: localities, and representation in provinces, ecogeographic landscapes, vegetation formations and protected areas. Gayana, 87(2), 108-122 - get paper here
  • Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here
  • Demangel, Diego 2016. Reptiles en Chile. Fauna Nativa Ediciones, Santiago, 619 pp - get paper here
  • Esquerré, Damien; J Scott Keogh, Diego Demangel, Mariana Morando, Luciano J Avila, Jack W Sites, Jr., Francisco Ferri-Yáñez, Adam D Leaché 2022. Rapid Radiation and Rampant Reticulation: Phylogenomics of South American Liolaemus Lizards. Systematic Biology 71 (2): 286–300 - get paper here
  • Lobo, F.; Espinoza, R.E. and S. Quinteros 2010. A critical review and systematic discussion of recent classification proposals for liolaemid lizards. Zootaxa 2549: 1–30 - get paper here
  • Marambio-Alfaro, Yery, Jaime Troncoso-Palacios 2014. Una nueva especie de Liolaemus del grupo de L. nigromaculatus (Iguania: Liolaemidae) para la Región de Atacama, Chile. Basic and Applied Herpetology 28: 65-77 - get paper here
  • Meiri, Shai; Aaron M. Bauer, Allen Allison, Fernando Castro-Herrera, Laurent Chirio, Guarino Colli, Indraneil Das, Tiffany M. Doan, Frank Glaw, Lee L. Grismer, Marinus Hoogmoed, Fred Kraus, Matthew LeBreton, Danny Meirte, Zoltán T. Nagy, Cristiano d 2017. Extinct, obscure or imaginary: the lizard species with the smallest ranges. Diversity and Distributions - get paper here
  • Núñez,H.; Schulte, J.A. & Garín, C. 2001. Liolaemus josephorum, nueva especie de lagartija para el Norte de Chile. Bol. Mus. Nac.Hist. Nat. 50: 91-107
  • Ortiz J C 1987. Une nouvelle espece de Liolaemus (Sauria, Iguanidae) du Chili. Bulletin du Muséum d’Histoire Naturelle SECTION A ZOOLOGIE BIOLOGIE ET ECOLOGIE ANIMALES 9(1) 1987: 265-270
  • Panzera A, Leaché AD, D’Elía G, Victoriano PF. 2017. Phylogenomic analysis of the Chilean clade of Liolaemus lizards (Squamata: Liolaemidae) based on sequence capture data. PeerJ 5:e3941 - get paper here
  • PINCHEIRA-DONOSO, Daniel & HERMAN NÚÑEZ 2005. The Chilean species of the genus Liolaemus Wiegmann, 1834 (Iguania, Tropiduridae, Liolaeminae). Taxonomy, systematics and evolution [in Spanish] Mus Nac Hist Nat Chile Publ Ocas 59: 1-486 - get paper here
  • Troncoso-Palacios, Jaime 2014. Nueva lista actualizada de los reptiles terrestres de la Región de Atacama, Chile. Boletín Chileno de Herpetología 1: 1-4 - get paper here
  • Troncoso-Palacios, Jaime & Francisco Ferri-Yánez 2012. REVISIÓN DEL ESTATUS TAXONÓMICO DE LIOLAEMUS JOSEPHORUM Núñez, Schulte & Garin 2001 (Iguania: Liolaemidae). BOLETIN DEL MUSEO REGIONAL DE ATACAMA (3): 93-101
 
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