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Liolaemus zullyae CEI & SCOLARO, 1996

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Higher TaxaLiolaemidae, Iguania, Sauria, Squamata (lizards)
Subspecies 
Common Names 
SynonymLiolaemus zullyi CEI & SCOLARO 1996: 393
Liolaemus zullyae — MICHELS & BAUER 2004
Liolaemus (Donosolaemus) zullyi — PINCHEIRA-DONOSO & NÚÑEZ 2005: 74
Liolaemus scolaroi PINCHEIRA-DONOSO & NUNEZ 2005: 70
Liolaemus zullyi — DUBOIS 2007 (by implication)
Liolaemus zullyi — PINCHEIRA-DONOSO et al. 2008
Liolaemus zullyi — LOBO et al. 2010
Liolaemus scolaroi — LOBO et al. 2010
Liolaemus scolaroi — AVILA et al. 2020
Liolaemus zullyae x Liolaemus scolaroi — SÁNCHEZ et al. 2023
Liolaemus zullyae — SÁNCHEZ et al. 2024 
DistributionArgentina (Santa Cruz Province), Chile

Type locality: "Rio Zeballos valley, in the Jeinement River basin, at 850 m elevation, in the neighbourhood of the ford of the river, lying on the track to Paso Roballos, some 80 km south of Los Antiguos, west of Perito Moreno, Santa Cruz Province, Argentina".

scolaroi: Chile, Argentina; Type locality: “Jeinimeni, XI Región Admisitrativa, Chile"  
Reproductionovoviviparous 
TypesHolotype: MRSN R-1141-1, collected by J.M. Cei and J.A. SCOLARO, 24 February 1995.
Holotype: MNHNC 3674 (Mus. Nac. Hist. Nat. Chile, Santiago) [scolaroi] 
Diagnosis 
CommentGroup: kingii group.

Hybridization: Liolaemus zullyae and Liolaemus scolaroi apparently hybridize in the wild fide Sánchez et al. 2024.

Synonymy: Demangel (2016) suggested that Liolaemus scolaroi is a synonym of Liolaemus zullyae, which was rejected and rebutted by Scolaro et al. 2018 but confirmed by Sanchez et al. 2024. L. archeforus is genetically indistinguishable from L. zullyi but morphologically clearly distinguishable, so they appear to be 2 morphs of the same species (Sánchez et al. 2023).

Distribution: Liolaemus scolaroi was erroneously omitted for Chile by Demangel 2016, but found there (D. Pincheira-Donoso, pers. comm., 23 May 2017).

NCBI taxonID: 971281 [scolaroi] 
EtymologyNamed after Mrs. Zully Ortega de Scolaro, the wife of the senior author, hence the name should be “zullyae”, not “zullyi”.

L. scolaroi: “Esta especie esta dedicada al Doctor José Alejandro Scolaro, por su importante contribución al conocimiento de la herpetofauna patagónica, en particular el área geográfica en que se distribuye la presente especie. Sea este un homenaje a un brillante herpetólogo argentino.” 
References
  • Avila, Luciano J.; Andrea González Marín, Jaime Troncoso-Palacios, Kevin Imanol Sánchez, Cristian Hernán Fulvio Pérez, and Mariana Morando 2020. Naming the Diversity: Taxonomy of Current Species of Patagonian Lizards. In: Morando & Avila (eds.), Lizards of Patagonia. Springer Nature, pp. 123-188 - get paper here
  • AVILA, LUCIANO JAVIER; LORENA ELIZABETH MARTINEZ & MARIANA MORANDO 2013. Checklist of lizards and amphisbaenians of Argentina: an update. Zootaxa 3616 (3): 201–238 - get paper here
  • Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here
  • Breitman, M. Florencia; , Luciano J. Avila, Jack W. Sites Jr., and Mariana Morando 2011. Lizards from the end of the world: Phylogenetic relationships of the Liolaemus lineomaculatus section (Squamata: Iguania: Liolaemini). Molecular Phylogenetics and Evolution 59 (2): 364-376 - get paper here
  • Breitman, María Florencia; Mariana Morando and Luciano Javier Avila 2013. Past and present taxonomy of the Liolaemus lineomaculatus section (Liolaemidae): is the morphological arrangement hypothesis valid? Zoological Journal of the Linnean Society 168 (3): 612–668 - get paper here
  • Breitman, María Florencia; Minoli, Ignacio; Avila, Luciano Javier; Medina, Cintia Debora; Sites, Jr., Jack Walter; Morando, Mariana 2014. Lagartijas de la provincia de Santa Cruz (Argentina): distribución geográfica, diversidad genética y estado de conservación . Cuad. herpetol. 28 (2): 83-110 - get paper here
  • Cei J M. Scolaro J A. 1996. A new species of Liolaemus of the archeforus group from the precordilleran valley of the Zeballos river, Santa Cruz Province, Argentina (Reptilia, Tropiduridae). Boll. Mus. Reg. Sci. Nat. Torino 14 (2): 389-401
  • Demangel, Diego 2016. Reptiles en Chile. Fauna Nativa Ediciones, Santiago, 619 pp - get paper here
  • Dubois, A. 2007. Genitives of species and subspecies nomina derived from personal names should not be emended. Zootaxa 1550: 49–68 - get paper here
  • Michels, J.P. & A.M. Bauer 2004. Some corrections to the scientific names of amphibians and reptiles. Bonner Zoologische Beiträge 52: 83–94 [2005] [Note: Dubois 2007 rejected most of the changes made in this paper] - get paper here
  • Minoli, Ignacio, Mónica L. Kozykariski, and Luciano J. Avila. 2010. Observations on parturition in two Liolaemus species of the archeforus group (Iguania: Squamata: Liolaemidae). Herpetology Notes 3: 333-336
  • PINCHEIRA-DONOSO, Daniel & HERMAN NÚÑEZ 2005. The Chilean species of the genus Liolaemus Wiegmann, 1834 (Iguania, Tropiduridae, Liolaeminae). Taxonomy, systematics and evolution [in Spanish] Mus Nac Hist Nat Chile Publ Ocas 59: 1-486 - get paper here
  • Ruiz-Monachesi, M.R., Valdecantos, S., Gamarra-Nallar, R., Moreno Azócar, D.L., Pueta, M., Perotti, G. and Cruz, F.B. 2024. Evolutionary novelty in chemical signalling is linked with chemosensory behaviour in close Liolaemus lizard species (Iguania: Liolaemidae). J Zool, 324: 253-266 - get paper here
  • Sánchez, Kevin I.; Hans Recknagel, Kathryn R. Elmer, Luciano J. Avila and Mariana Morando 2024. Tracing evolutionary trajectories in the presence of gene flow in South American temperate lizards (Squamata: Liolaemus kingii group). Evolution qpae009 - get paper here
  • Sánchez, Kevin I; Emilce G Diaz Huesa, María F Breitman, Luciano J Avila, Jack W Sites Jr., Mariana Morando 2023. Complex Patterns of Diversification in the Gray Zone of Speciation: Model-Based Approaches Applied to Patagonian Liolaemid Lizards (Squamata: Liolaemus kingii clade). Systematic Biology syad019 - get paper here
 
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