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Lipinia inexpectata DAS & AUSTIN, 2007

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Higher TaxaScincidae, Sphenomorphinae (Sphenomorphini), Scincoidea, Sauria, Squamata (lizards)
Common Names 
SynonymLipinia inexpectata DAS & AUSTIN 2007
Saiphos quadrivittatum — DE ROOIJ 1915: 271
Saiphos quadrivittatum — TAYLOR 1963: 1053
Lipinia quadrivittata — GREER 1974: 11
Lipinia quadrivittata — BROWN & ALCALA 1980: 85
Lipinia sp. — DAS 2004 
DistributionIndonesia (Borneo, Celebes)

Type locality: Pulau Manukan (05°58’N; 115°59’E), Kota Kinabalu District, Sabah, East Malaysia  
Reproductionoviparous (not imputed, fide Zimin et al. 2022) 
TypesHolotype: ZRC 2.5728 (field number ID– 7191; holotype); C. C. Austin and I. Das, collectors, 5 December 2000. Adult female (with paired shelled ova in the oviducts). 
DiagnosisDiagnosis: A small (SVL to 40.6 mm) species of Lipinia, diagnosable from congeneric species in having the following combination of characters: external ear opening absent; lower eyelid with a clear spectacle; 20 midbody scale rows; 46–50 longitudinal scale rows between parietals and base of tail; 16–17 lamellae under toe IV; 6 supralabials; 6–7 infralabials; 68–74 subcaudals; dorsum tan brown with dark gray brown stripes; dorsum tan brown with a series of dark gray brown stripes; a series of dark spots arranged in a longitudinal series bilaterally on each outer side of the paravertebral stripes; labials unbarred; venter yellowish-cream with pale gray smudges, and a unique cytb gene sequence. 
CommentPopulations of L. inexpectata from Borneo have been previously described as L. quadrivittata. 
EtymologyThe species name (an adjective in feminine form) refers to the unexpected nature of the discovery of the type series. This species was referred to Lipinia quadrivittata for 90 years since the work of de Rooij (1915). 
  • Brown, WC. & A.C. ALCALA 1980. Philippine Lizards of the family Scincidae. Silliman Univ. Nat. Sci., Dumaguete City, Mon., Ser. 2: i-xi + 1-246.
  • Das , I. 2004. Lizards of Borneo. Natural History Publications, Kota Kinabalu, Borneo
  • Das, I. & Austin, C.C. 2007. New Species of Lipinia (Squamata: Scincidae) from Borneo, Revealed by Molecular and Morphological Data. Journal of Herpetology 41 (1): 61 - get paper here
  • Greer, A.E. 1973. Two new lygosomine skinks from New Guinea with comments on the loss of the external ear in lygosomines and observations on previously described species. Breviora (406): 1-25 - get paper here
  • Taylor, E.H. 1963. The lizards of Thailand. Univ. Kansas Sci. Bull. 44: 687-1077. - get paper here
  • Zimin, A., Zimin, S. V., Shine, R., Avila, L., Bauer, A., Böhm, M., Brown, R., Barki, G., de Oliveira Caetano, G. H., Castro Herrera, F., Chapple, D. G., Chirio, L., Colli, G. R., Doan, T. M., Glaw, F., Grismer, L. L., Itescu, Y., Kraus, F., LeBreton 2022. A global analysis of viviparity in squamates highlights its prevalence in cold climates. Global Ecology and Biogeography, 00, 1–16 - get paper here
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