Lophocalotes achlios HARVEY, SCRIVANI, SHANEY, HAMIDY, KURNIAWAN & SMITH, 2018

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Higher Taxa	Agamidae (Draconinae), Sauria, Iguania, Squamata (lizards)
Subspecies
Common Names	E: White-throated Crested Dragon
Synonym	Lophocalotes achlios HARVEY, SCRIVANI, SHANEY, HAMIDY, KURNIAWAN & SMITH 2018
Distribution	Indonesia (Sumatra) Type locality: along the main trail up Gunung Kaba, Kabupaten Rejang Lebong, Provinsi Bengkulu, Sumatera, Indonesia, 3.50757°S, 102.62927°E, 1818 m elevation (a.s.l.).
Reproduction	oviparous (not imputed, fide Zimin et al. 2022)
Types	Holotype: MZB 14038, adult male, field tag ENS 18170; Figs. 1, 2A,B) collected 16 July 2015 by M.B. Harvey, E. Wostl, G.C. Sarker, and P. Thammachoti. Paratypes (n = 20). Eleven males (MZB 14030, 14032, 14033, 14037–14039, UTA 63730, 63733, 63735–63737) and eight females (MZB 14031, 14034–14036, UTA 63729, 63731, 63732, 63734) collected on 16 July 2015 by M.B. Harvey, E. Wostl, G.C. Sarker, and P. Thammachoti along the main trail up Gunung Kaba, 3.50026–3.50757°S, 102.62927–102.63557°E, 1567–1818 m a.s.l.; and one male (UTA 63740) collected by E.N. Smith, E. Wostl, G.C. Sarker, and G. Pradana on 22 July 2015 from a trail up Gunung Kambing above the police academy near the road to Gunung Kaba, 3.39649°S, 102.63599°E, 1636 m a.s.l., Kabupaten Rejang Lebong, Provinsi Bengkulu, Sumatera, Indonesia.
Diagnosis	Diagnosis: A species of Lophocalotes differing from the L. ludekingi in having more gulars, ventrals, and subdigital lamellae; in having males with a lower nuchal crest not supported by an arched flap of skin and white gular markings; and in having females with cream buccal epithelia.
Comment
Etymology	The specific epithet achlios is a feminine noun in apposition derived by affixing the Greek suffix –ios meaning ‘‘pertaining to’’ or ‘‘of’’ to achlys, meaning mist. The new name refers to the misty cloud forests where Lophocalotes achlios occurs.
References	Harvey, Michael B.; James Scrivani, Kyle Shaney, Amir Hamidy, Nia Kurniawan, and Eric N. Smith 2018. Sumatra's Endemic Crested Dragons (Agamidae: Lophocalotes): A New Species from the Bukit Barisan Range, Comments on Lophocalotes ludekingi, and Ecology. Herpetologica 74 (1): 73-88. - get paper here Kwet, A. 2019. Liste der im Jahr 2018 neu beschriebenen Reptilien. Elaphe 2019 (3): 52-72 Zimin, A., Zimin, S. V., Shine, R., Avila, L., Bauer, A., Böhm, M., Brown, R., Barki, G., de Oliveira Caetano, G. H., Castro Herrera, F., Chapple, D. G., Chirio, L., Colli, G. R., Doan, T. M., Glaw, F., Grismer, L. L., Itescu, Y., Kraus, F., LeBreton 2022. A global analysis of viviparity in squamates highlights its prevalence in cold climates. Global Ecology and Biogeography, 00, 1–16 - get paper here
External links	IUCN Red List National Center for Biotechnology Information Google images

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