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Lubuya ivensii (BOCAGE, 1879)

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Higher TaxaScincidae, Mabuyinae (Mabuyini), Scincoidea, Sauria, Squamata (lizards)
Common NamesE: Iven’s Water Skink, Meadow Skink, Iven’s skink 
SynonymEuprepes ivensii BOCAGE 1879
Mabuia ivensii — BOULENGER 1887: 197
Lygosoma ivensii — BOCAGE 1895
Mabuya ivensii — MANACAS 1963
Mabuya ivensi septemlineata LAURENT 1964
Lubuya ivensii — HORTON 1972
Mabuya ivensii — GREER et al. 2000
Euprepis ivensii — MAUSFELD et al. 2002
Trachylepis ivensii — BAUER 2003
Lubuya ivensii — CONRADIE et al. 2016
Labuya ivensii — WEINELL et al. 2019 (in error) 
DistributionAngola, NW Zambia, S Democratic Republic of the Congo (Zaire)

Type locality: interior of Benguella, Angola; restricted to “Benguela” by BOULENGER 1887.  
ReproductionOvoviviparous. Females ovulate tiny eggs and later supply the nutrients for development by placental means. 
TypesHolotype: lost, was MB (Museu Bocage) (probably lost in fire 1978)
Types: MD (Museu Dundo) [septemlineata] 
DiagnosisDiagnosis (genus): Related to Mabuya, from which it is readily distinguishable by the position of the nostrils on top of the snout, the elongation of the postnasal scale and the extreme elongation of the body (from Horton 1972). 
CommentSynonymy: partly after Horton 1972. Branch and Haagner (1993) synonymized Lubuya with Mabuya but Metallinou et al. 2016 revived it again.

Type species: Euprepes ivensii BOCAGE 1879 is the type species of the genus Lubuya HORTON 1972. 
EtymologyNamed after the collector of the type specimens, Roberto Ivens (b. 12 June 1850, Ponta Delgada, Portugal, d. 28 January 1898, Cruz Quebrada - Dafundo, Portugal). (Glenn Shea, pers. comm., 2 Feb 2024) 
  • Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here
  • Blackburn, D.G. & Flemming, A.F. 2011. Invasive implantation and intimate placental associations in a placentotrophic african lizard, Trachylepis ivensi (scincidae). Journal of Morphology 273: 137–159. doi: 10.1002/jmor.11011 - get paper here
  • Blackburn, Daniel G. and Alexander F. Flemming 2010. Reproductive specialization in a viviparous African skink and its implications for evolution and conservation. Herp. Cons. Biol. 5 (2) - get paper here
  • Bocage,J.V. du B. 1895. Herpétologie d'Angola et du Congo. Lisbon: Imprimerie Nationale, i-xx, 203 pp. - get paper here
  • Bocage,J.V.B. du 1879. Reptiles et batraciens nouveaux d'Angola. J. Acad. Sci. Lisbon 7: 97-99
  • Boulenger, G. A. 1887. Catalogue of the lizards in the British Museum (Nat. Hist.) III. Lacertidae, Gerrhosauridae, Scincidae, Anelytropsidae, Dibamidae, Chamaeleontidae. London: 575 pp. - get paper here
  • Branch W. R.; Haagner, G.V. 1993. The skink Mabuya ivensii: new records from Zambia and Zaire, and the status of the subspecies septemlineata Laurent 1964 and the genus Lubuya Horton. Amphibia-Reptilia 14 (2): 105-115 - get paper here
  • Broadley, D.G. 1998. The reptilian fauna of the Democratic Republic of the Congo (Congo-Kinshasa). In: Schmidt, K.P. and Noble, G.K., Contributions to the Herpetology of the Belgian Congo... [reprint of the 1919 and 1923 papers]. SSAR Facsimile reprints in Herpetology, 780 pp.
  • Broadley, Donald G. and Frank Willems. 2015. Geographic Distribution. Trachylepis ivensii Bocage, 1879, Meadow Skink. African Herp News (62): 41-42 - get paper here
  • Ceríaco, Luis M. P.; Mariana P. Marques, Ilunga André, Esteves Afonso, David C. Blackburn & Aaron M. Bauer 2020. ILLUSTRATED TYPE CATALOGUE OF THE “LOST” HERPETOLOGICAL COLLECTIONS OF MUSEU DO DUNDO, ANGOLA. Bulletin of the Museum of Comparative Zoology 162 (7): 379-440 - get paper here
  • Conradie W, Keates C, Verburgt L, Baptista NL, Harvey J, Júlio T, Neef G. 2022. Contributions to the herpetofauna of the Angolan Okavango-Cuando-Zambezi River drainages. Part 2: Lizards (Sauria), chelonians, and crocodiles. Amphibian & Reptile Conservation 16(2): 181–214 (e322) - get paper here
  • Greer, A.E., Arnold, C. & Arnold, E.N. 2000. The systematic significance of the number of presacral vertebrae in the scincid lizard genus Mabuya. Amphibia-Reptilia 21 (1): 121-126 - get paper here
  • Haagner,G.V.; Branch,W.R. & Haagner,A.J.F. 2000. Notes on a collection of reptiles from Zambia and adjacent areas of the Democratic Republic of the Congo. Annals of the Eastern Cape Museum 1: 1 – 25
  • Horton, D. R. 1972. A new scincid genus from Angola. Journal of Herpetology 6 (1): 17-20 - get paper here
  • Laurent, R.F. 1964. Reptiles et batraciens de l'Angola (troisième contribution). Companhia de Di amantes de Angola (Diamang), Serviços Culturais, Museu do Dundo (Angola), No. 67, 165 pp.
  • Manacas,S. 1963. Saurios de Angola. Mem. Junta Invest. Ultram. 2nd ser. 43: 223-240
  • Marques, Mariana P.; Luis M. P. Ceríaco , David C. Blackburn , and Aaron M. Bauer 2018. Diversity and Distribution of the Amphibians and Terrestrial Reptiles of Angola -- Atlas of Historical and Bibliographic Records (1840–2017). Proc. Cal. Acad. Sci. (Ser. 4) 65: 1-501 (Supplement II)
  • Pietersen, Darren, Verburgt, Luke & Davies, John 2021. Snakes and other reptiles of Zambia and Malawi. Struik Nature / Penguin Random House South Africa, 376 pp., ISBN 9781775847373
  • Wagner, Philipp; Dennis Rödder & Thomas M. Wilms 2012. New data on the morphology and natural history of Tetradactylus ellenbergeri (Angel, 1922) (Sauria: Gerrhosauridae) and Trachylepis ivensii (Bocage, 1879) (Sauria: Scincidae) in northeastern Zambia. Bonn zoological Bulletin 61 (1): 35–40 - get paper here
  • Weinell, J. L., Branch, W. R., Colston, T. J., Jackman, T. R., Kuhn, A., Conradie, W., & Bauer, A. M. 2019. A species-level phylogeny of Trachylepis (Scincidae: Mabuyinae) provides insight into their reproductive mode evolution. Molecular Phylogenetics and Evolution - get paper here
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