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Lucasium damaeum (LUCAS & FROST, 1896)

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Higher TaxaDiplodactylidae, Gekkota, Sauria, Squamata (lizards: geckos)
Common NamesE: Beaded Gecko 
SynonymCeramodactylus damaeus LUCAS & FROST 1896
Lucasius damaeus — KINGHORN 1929: 77
Lucasius damaeus — LOVERIDGE 1934: 297
Lucasium damaeum — WERMUTH 1965: 100 (nom. nov.)
Diplodactylus damaeus — KLUGE 1967
Diplodactylus daemeus — COVENTRY 1970
Oedura lesueurii lesueurii — DAVEY 1970 (nec O. lesueurii DUMÉRIL & BIBRON)
Diplodactylus dameum — GORMAN 1973
Turnerdactylus damaeus — WELLS & WELLINGTON 1989
Diplodactylus damaeus — KLUGE 1993
Diplodactylus damaeus — RÖSLER 1995: 80
Lucasium damaeum — COGGER 2000: 250
Diplodactylus damaeus — RÖSLER 2000: 77
Diplodactylus damaeus — GREER 2006 (online)
Lucasium damaeum — OLIVER et al. 2007 
DistributionAustralia (drier interior from SE Western Australia to S Queensland and W New South Wales and Victoria)

Type locality: Charlotte Waters, Northern Territory.  
TypesHolotype: NMV D7528 (formerly 57846) 
DiagnosisDiagnosis (genus): OLIVER 2007.

L. damaeum is characterized by complete loss of subdigital lamellae and scansors, but Kluge (1967) suggested that this species was no more than the extreme of a trend seen in the stenodactylus group of Diplodactylus.

Description. Head large, high ; snout obtusely pointed, a little longer than the distance between the orbit and the ear opening. Ear-opening narrow, elliptical, oblique. Body slightly depressed. Limbs moderate, the fore-limb stretched forward reaches to between the eye and the nostril. Digits long, slender, inferiorly with small, imbricate, pointed scales. Head and upper surfaces of body, and limbs, covered with small granular scales. Rostral quadrangular, twice as broad as high, with median cleft above. Nostril pierced between the rostral, first labial, and four nasals, the supero-anterior nasals large, forming a suture with one another behind the rostral. Eleven or twelve upper and as many lower labials. Mental rather large, trapezoid ; no chin shields. Gular scales very small, granular. Abdominal scales flat, subimbricate. Male with two or three blunt spines on each side of the base of the tail, and two widely separated præanal pores. Tail missing. Colour.-Pale whitish-grey above, darkest on the sides; a brownish, more or less broken band from the snout along each side of the back to tail; a broad, median whitish band from neck to base of tail head spotted oL reticulated with dark brown; sides with two longitudinal series of roundish white spots limbs and under surfaces uniform whitish. (Lucas & Frost 1896: 1) 
CommentSynonymy: Assigned previously to the Diplodactylus stenodactylus group. The name Lucasius is preoccupied by a Crustacean and has also been used for a Hymenopteran). WELLS & WELLINGTON 1984 synonymized Lucasium with Diplodactylus. However, Lucasium was resurrected by Oliver et al. 2007. McGrath 2008 suggested to merge Lucasium and Diplodactylus into the genus Reliquodactylus, an unnecessary act that produced nothing but a new synonym and actually a nomen nudum (Rösler et al. 2018, forthcoming).

Type species: Ceramodactylus damaeus is the type species of the genus Lucasium WERMUTH 1965. 
EtymologyThe genus Lucasium has been named after Arthur Henry Shakespeare LUCAS (1853-1936), british-born physician and herpetologist who moved to Australia in 1883. For more biographical details see Shea 2003. 
  • CHIACCHIO, MICHELE; ANNEGRET GRIMM-SEYFARTH<br />KLAUS HENLE<br /> AND JEAN-BAPTISTE MIHOUB 2020. Water availability as a major climatic driver of taxonomic and functional diversity in a desert reptile community. Ecosphere 11(7):e03190. 10.1002/ecs2.3190 - get paper here
  • Cogger, H. G. 2014. Reptiles and Amphibians of Australia, 7th ed. CSIRO Publishing, xxx + 1033 pp. - get paper here
  • Cogger, H.G. 2000. Reptiles and Amphibians of Australia, 6th ed. Ralph Curtis Publishing, Sanibel Island, 808 pp.
  • Ferguson, Daniel J.; Mathieson, Michael T.; Eyre, Teresa J. 2015. ADDITIONAL INFORMATION ON THE BEADED GECKO Lucasium damaeum AND SMOOTH KNOB-TAILED GECKO Nephrurus levis AT THEIR EASTERN LIMIT IN SOUTHWEST QUEENSLAND. Queensland Naturalist 53 (1-3): 19-28 - get paper here
  • Kinghorn, J. Roy 1924. Reptiles and batrachians from south and south-west Australia. Rec. Austral. Mus. 14 (3): 163-183 - get paper here
  • Kinghorn, J. Roy 1929. Herpetological notes 1. Rec. Austral. Mus. 17 (2): 76-84 - get paper here
  • Loveridge, A. 1934. Australian reptiles in the Museum of Comparative Zoology, Cambridge, Massachusetts. Bull. Mus. Comp. Zool. Harvard 77: 243-383 - get paper here
  • Lucas, A. H. S., and C. Frost. 1896. Further preliminary notice of certain new species of lizards from central Australia. Proceedings of the Royal Society of Victoria, 8:1-4. - get paper here
  • McGrath, J. 2008. Geckos, Family Gekkonidae. In: Swan M. (Ed), Keeping & Breeding Australian Lizards. Mike Swan Herp Books, Lilydale, Australia, pp. 73–213
  • Oliver, Paul M.; Hutchinson, Mark N.; Cooper, Steven J.B. 2007. Phylogenetic relationships in the lizard genus Diplodactylus Gray and resurrection of Lucasium Wermuth (Gekkota, Diplodactylidae). Australian Journal of Zoology 55( 3):197-210 - get paper here
  • Pepper, Mitzy; Paul Doughty, J. Scott Keogh 2006. Molecular phylogeny and phylogeography of the Australian Diplodactylus stenodactylus (Gekkota; Reptilia) species-group based on mitochondrial and nuclear genes reveals an ancient split between Pilbara and non-Pilbara D. stenodactylus. Molecular Phylogenetics and Evolution 41: 539–555 - get paper here
  • Rösler, H. 2000. Kommentierte Liste der rezent, subrezent und fossil bekannten Geckotaxa (Reptilia: Gekkonomorpha). Gekkota 2: 28-153
  • Rösler, Herbert 1995. Geckos der Welt - Alle Gattungen. Urania, Leipzig, 256 pp.
  • Schönecker, P. 2007. Biologie, Haltung und Nachzucht des australischen Bodengeckos Lucasium damaeum. Draco 8 (29): 31-38 - get paper here
  • Shea, G.M. 2003. The HORN Expedition (1894) to Central Australia: New Directions in Australian Herpetology. Bonner Zoologische Beiträge 52 (3/4): 245-273 - get paper here
  • Swan, G.; Sadlier, R.; Shea, G. 2017. A field guide to reptiles of New South Wales. Reed New Holland, 328 pp.
  • Wilson, S. & Swan, G. 2010. A complete guide to reptiles of Australia, 3rd ed. Chatswood: New Holland, 558 pp.
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