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Lycodon subannulatus (DUMÉRIL, BIBRON & DUMÉRIL, 1854)

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Higher TaxaColubridae, Colubrinae, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies 
Common NamesE: Malayan Bridal Snake, Southern Bridle Snake
G: Südliche Zügelnatter 
SynonymOdontomus sub-annulatus DUMÉRIL, BIBRON & DUMÉRIL 1854: 454
Odontomus subannulatus — BLEEKER 1858
Odontomus subannulata — JAN 1863
Nymphodidium subannulatus — BLANFORD 1881
Hydrophobus subannulatus — BOULENGER 1890
Dryocalamus subannulatus — BOULENGER 1893: 371
Dryocalamus cf. subannulatus — MALKMUS & KUNKEL 1992
Nymphodidium maculatum GÜNTHER 1864
Nymphophidium maculatum — MÜLLER 1880
Coronella prosopeion BLEEKER (nomen nudum)
Dryocalamus davisonii tungsongensis NUTPHAND 1986
Dryocalamus subannulatus — MANTHEY & GROSSMANN 1997: 340
Dryocalamus subannulatus — COX et al. 1998: 65
Dryocalamus subannulatus — WALLACH et al. 2014: 248
Dryocalamus subnanulatus — KAVIARASU et al. 2014
Lycodon subannulatus — FIGUEROA et al. 2016
Dryocalamus subnanulatus — DE LANG 2017
Dryocalamus subnanulatus — LEE et al. 2018
Dryocalamus subannulatus — WICKRAMASINGHE et al. 2020 
DistributionIndonesia (Mentawai Archipelago, Riau Archipelago, Sumatra, Java, Natuna islands); Malaysia (Malaya and East Malaysia, Pulau Pangkor, Pulau Tioman, Johor: Pulau Sibu); Borneo
Philippines (Palawan);
Singapore; S Thailand, Myanmar (Thayawthadangyi Island group; Daung [Don] Island)

Type locality: Padang, West Sumatra  
Reproductionoviparous 
TypesType: BMNH 1946.1.13.80 (and possibly additional specimens). 
Diagnosis 
CommentVariation: there is also a lineated phase of Dryocalamus subannulatus.

Habitat: partly arboreal (Harrington et al. 2018). 
References
  • Boulenger, G.A. 1893. Catalogue of the snakes in the British Museum (Nat. Hist.) I. London (Taylor & Francis), 448 pp. - get paper here
  • Chan-ard, T., Parr, J.W.K. & Nabhitabhata, J. 2015. A field guide to the reptiles of Thailand. Oxford University Press, NY, 352 pp. [see book reviews by Pauwels & Grismer 2015 and Hikida 2015 for corrections] - get paper here
  • Cox, Merel J.; Van Dijk, Peter Paul; Jarujin Nabhitabhata & Thirakhupt,Kumthorn 1998. A Photographic Guide to Snakes and Other Reptiles of Peninsular Malaysia, Singapore and Thailand. Ralph Curtis Publishing, 144 pp.
  • Das, I. 2012. A Naturalist's Guide to the Snakes of South-East Asia: Malaysia, Singapore, Thailand, Myanmar, Borneo, Sumatra, Java and Bali. Oxford J, ohn Beaufoy Publishing - get paper here
  • David,P. & Vogel, G. 1996. The snakes of Sumatra. An annotated checklist and key with natural history notes. Bücher Kreth, Frankfurt/M.
  • Duméril, A. M.C., G. BIBRON & A. DUMÉRIL 1854. Erpétologie générale ou Histoire Naturelle complète des Reptiles. Vol. 9. Paris, XX + 440 S. - get paper here
  • Figueroa A, McKelvy AD, Grismer LL, Bell CD, Lailvaux SP 2016. A Species-Level Phylogeny of Extant Snakes with Description of a New Colubrid Subfamily and Genus. PLoS One 11 (9): e0161070.doi:10.1371/journal.pone.0161070 - get paper here
  • FIGUEROA, ALEX; MARTYN E.Y. LOW, KELVIN K.P. LIM 2023. Singapore’s herpetofauna: updated and annotated checklist, history, conservation, and distribution. Zootaxa 5287 (1): 001–378 - get paper here
  • Grismer, L.L. 2011. Amphibians and reptiles of the Seribuat Archipelago. Edition Chimaira, Frankfurt, 239 pp.
  • Grismer, L.L., McGuire, J.A.; Sosa, R. & Kaiser, H. 2002. Revised checklist and comments on the terrestrial herpetofauna of Pulau Tioman, Peninsular Malaysia. Herpetological Review 33 (1): 26-29 - get paper here
  • Harrington, Sean M; Jordyn M de Haan, Lindsey Shapiro, Sara Ruane 2018. Habits and characteristics of arboreal snakes worldwide: arboreality constrains body size but does not affect lineage diversification. Biological Journal of the Linnean Society 125 (1): 61–71 - get paper here
  • Herlambang, A. E., Riyanto, A., Munir, M., Hamidy, A., Kimura, K., Eto, K., & Mumpuni, M. 2022. AFTER 16 YEARS: AN UPDATE CHECKLIST OF HERPETOFAUNA ON THE NATUNA ISLANDS, INDONESIA. TREUBIA, 49(2), 67-84 - get paper here
  • Janssen HY, Pham CT, Ngo HT, Le MD, Nguyen TQ, Ziegler T 2019. A new species of Lycodon Boie, 1826 (Serpentes, Colubridae) from northern Vietnam. ZooKeys 875: 1-29 - get paper here
  • Karraker, Nancy E., Colin T. Strine, Matthew Carne and Anne Devan-Song. 2015. Dryocalamus subannulatus (Malayan bridle snake) behavior. Herpetological Review 46 (2): 272-273 - get paper here
  • Kaviarasu, M; MA Shahfiz, MR Rizal, RT John, FMR Nordiana & IF Fakrul 2014. ADDITION TO THE HERPETOFAUNA DIVERSITY IN THE FOREST RESEARCH INSTITUTE MALAYSIA (FRIM) CAMPUS, SELANGOR. PROCEEDINGS OF THE CONFERENCE ON FORESTRY AND FOREST PRODUCTS RESEARCH 2013, Kala Lumpur, pp. 324-329
  • Lang, Ruud de 2017. The Snakes of Java, Bali and Surrounding Islands. Edition Chimaira, Frankfurt am Main, Germany, 435 pp. - get paper here
  • Lee JL, Miller AH, Connette GM, Oo KS, Zug GR, Mulcahy DG 2018. First record of the Malaysian Bridle Snake, Dryocalamus subannulatus (Duméril, Bibron & Duméril, 1854), in Myanmar (Reptilia, Serpentes, Colubridae). Check List 14(2): 341-345 [erratum in Check List 16(5): 1393-1393, 2020] - get paper here
  • Leo S, Suherman M, Permatasari A, Suganda D, Zulamri, Winarni NL. 2020. Herpetofauna diversity in Zamrud National Park, Indonesia: baseline checklist for a Sumatra peat swamp forest ecosystem. Amphibian & Reptile Conservation 14(2) [General Section]: 250–263 (e249) - get paper here
  • Leong, Tzi Ming 2001. Geographic distribution. Dryocalamus subannulatus. Herpetological Review 32 (3):195 - get paper here
  • Manthey, U. & Grossmann, W. 1997. Amphibien & Reptilien Südostasiens. Natur und Tier Verlag (Münster), 512 pp. - get paper here
  • Müller, F. 1880. Erster Nachtrag zum Katalog der herpetologischen Sammlung des Basler Museums. Verh. Naturf. Ges. Basel 7: 120-165 - get paper here
  • Onn, Chan Kin; J. van Rooijen, L. Lee Grismer, Daicus Belabut,, Mohd. Abdul Muin Md. Akil, Hamidi Jamaludin, Rick Gregory, and Norhayati Ahmad 2010. FIRST REPORT ON THE HERPETOFAUNA OF PULAU PANGKOR, PERAK, MALAYSIA. Russ. J. Herpetol. 17 (2): 139 – 146 - get paper here
  • Pauwels, O.S.G.; David, P. & Chan-ard, T. 2006. Dryocalamus davisonii tungsongensis Nutphand, 1986 and Lycodon suratensis Nutphand, 1986 (Serpentes: Colubridae): translation of their original description and taxonomic status. Hamadryad 30 (1-2):114 – 120 - get paper here
  • Pauwels,O.S.G. et al. 2000. Herpetological investigations in Phang-Nga Province, southern Peninsular Thailand, with a list of reptile species and notes on their biology. Dumerilia 4 (2): 123-154 - get paper here
  • Smedley, N. 1931. Notes on some Malaysian snakes. Bull. Raffles Mus. No 5: 49-54 - get paper here
  • Stuebing, R.B., Inger, R.F. & Lardner, B. 2014. A field guide to the snakes of Borneo, second edition. Natural history Publications (Borneo)
  • Syafiq MF, Badli-Sham BH, Grismer LL, Ahmad AB 2023. Uneven species occurrence and richness of lowland snakes (Serpentes, Squamata) in Terengganu, Peninsular Malaysia, with new locality records. ZooKeys 1168: 11–39
  • Teo, R.C.H. & N.J. Thomas 2019. Updated Inventory of Amphibians, Reptiles and Mammals of the Bukit Timah Nature Reserve, Singapore. Gardens’ Bulletin Singapore 71(Suppl. 1):145-183; doi: 10.26492/gbs71(suppl.1).2019-06 - get paper here
  • Teo, R.C.H. & Rajathurai, S. 1997. Mammals, reptiles and amphibians in the Nature Reserves of Singapore - diversity, abundance and distribution. Proc. Nature Reserves Survey Seminar. Gardens’ Bulletin Singapore 49: 353-425
  • TEYNIÉ, ALEXANDRE; PATRICK DAVID, & ANNEMARIE OHLER 2010. Note on a collection of Amphibians and Reptiles from Western Sumatra (Indonesia), with the description of a new species of the genus Bufo. Zootaxa 2416: 1–43 - get paper here
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp.
  • Wood, Perry L.; Timothy M. Youmans, Jesse L. Grismer, Jonathan Wheatley, Steven Wright, Cindy Valdivia, Armando Ponce, Linda Escobar, Suzannah Amin, Porshia Baker, Joanna Bernard, Stephany Looper, Nicholas Marsh, Linette Martin, Nicole Padilla, Rober 2004. First report of the herpetofauna of Pulau Sibu, Johor, West Malaysia. Hamadryad 28 (1&2): 116-119 - get paper here
 
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