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Lycodon albofuscus (DUMÉRIL, BIBRON & DUMÉRIL, 1854)

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Higher TaxaColubridae, Colubrinae, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies 
Common NamesE: Dark Wolf Snake
G: Dunkle Wolfszahnnatter 
SynonymSphecodes albo-fuscus DUMÉRIL, BIBRON & DUMÉRIL 1854: 394
Sphecodes albo-fuscus DUMÉRIL 1853 (nom. nud., generic name not latinized)
Sphecodes albofuscus — BLEEKER 1858
Ophites albofuscus — GÜNTHER 1858
Ophites albofuscus — BOULENGER 1885: 389
Lycodon albofuscus — BOULENGER 1893: 357
Lepturophis borneensis BOULENGER 1900: 183
Lepturophis borneensis — GRANDISON 1972: 87
Lepturophis borneensis — GRANDISON 1978
Lycodon albofuscus — MANTHEY 1983
Lepturophis borneensis — MURPHY et al. 1994
Lepturophis albofuscus — MANTHEY & GROSSMANN 1997: 359
Lepturophis borneensis — COX et al. 1998: 64
Lepturophis albofuscus — CHAN-ARD et al. 1999: 169
Lycodon albofuscus — LANZA 1999
Lepturophis borneensis — STUEBING & INGER 1999
Lepturophis albofuscus — HIEN et al. 2001
Lepturophis albofuscus — GRISMER et al. 2002
Lycodon albofuscus — DAS & YAAKOB 2007
Lycodon albofuscus — DAS 2012
Lepturophis albofuscus — WALLACH et al. 2014: 370
Lepturophis albofuscus — STUEBING et al. 2014
Lycodon albofuscus — FIGUEROA et al. 2016 
DistributionIndonesia (Sumatra, Nias); Thailand (Narathiwat, Satun, Songkhla, Surat Thani, Nakhon Si Thammarat, Yala),
Malaysia (Sabah, Sarawak, Borneo), Pulau Tioman.

Type locality: Sumatra

borneensis: Indonesia (Nias, Sumatra, Borneo);
Malaysia (Malaya and East Malaysia; Sabah, Sarawak, Selangor, Pahang, Jahore); S Thailand; Type locality: Sarawak. Emended to Kuching, Sarawak by de Rooij 1917).  
Reproductionoviparous. 
TypesHolotype: lost fide Iskandar & Colijn, 2001: 65. Not located at MNHN (I. Ineich, March 2019) 
Diagnosis 
CommentJuveniles have white bands on the black body while adults are completely black.

Type species: Sphecodes albo-fuscus DUMÉRIL, BIBRON & DUMÉRIL 1854: 394 is the type species of the genus Lepturophis BOULENGER 1900.

Synonymy: Dryocalamus is currently (2017) nested within Lycodon (e.g. see phylogenetic tree in Wostl et al. 2017), hence the Philippine Lycodon must be moved to a new genus or Lepturophis synonymized with Lycodon.

Habitat: fully arboreal (Harrington et al. 2018). 
References
  • Boulenger, G.A. 1893. Catalogue of the snakes in the British Museum (Nat. Hist.) I. London (Taylor & Francis), 448 pp. - get paper here
  • Boulenger, G.A. 1900. Description of new reptiles and batrachians from Bomeo. Proc. Zool. Soc. London: 182-187. - get paper here
  • Boulenger,G.A. 1885. A list of reptiles and batrachians from the Island of Nias. Ann. Mag. nat. Hist. (5) 16: 388-389 - get paper here
  • Chan-ard, T., Parr, J.W.K. & Nabhitabhata, J. 2015. A field guide to the reptiles of Thailand. Oxford University Press, NY, 352 pp. [see book reviews by Pauwels & Grismer 2015 and Hikida 2015 for corrections] - get paper here
  • Chan-ard,T.; Grossmann,W.; Gumprecht,A. & Schulz,K. D. 1999. Amphibians and reptiles of peninsular Malaysia and Thailand - an illustrated checklist [bilingual English and German]. Bushmaster Publications, Würselen, Gemany, 240 pp. [book review in Russ. J Herp. 7: 87] - get paper here
  • Cota, Michael; Robert W. Mendyk , Tanya Chan-ard , Sunchai Makchai and Pratheep Meewattana 2011. Range Extension, New Records and Natural History Notes on Lepturophis albofuscus (Duméril, Bibron & Duméril, 1854) (Squamata: Colubridae) in Thailand. The Thailand Natural History Museum Journal 5(2): 151-153
  • Cox, Merel J.; Van Dijk, Peter Paul; Jarujin Nabhitabhata & Thirakhupt,Kumthorn 1998. A Photographic Guide to Snakes and Other Reptiles of Peninsular Malaysia, Singapore and Thailand. Ralph Curtis Publishing, 144 pp.
  • Das, I. 2012. A Naturalist's Guide to the Snakes of South-East Asia: Malaysia, Singapore, Thailand, Myanmar, Borneo, Sumatra, Java and Bali. Oxford J, ohn Beaufoy Publishing - get paper here
  • Das, I. & Yaakob, N. 2007. Status of knowledge of the Malaysian herpetofauna. In Status of biological diversity in Malaysia & threat assessment of plant species in Malaysia. in: L. S. L. Chua, L. G. Kirton & L. G. Saw (eds.), Status of biological diversity in Malaysia & threat assessment of plant species in Malaysia. Forest Research Institute Malaysia, Kepong, pp. 31-81
  • DAVID, Patrick; MEREL J. COX, OLIVIER S. G. PAUWELS, LAWAN CHANHOME AND KUMTHORN THIRAKHUPT 2004. Book Review - When a bookreview is not sufficient to say all: an in-depth analysis of a recent book on the snakes of Thailand, with an updated checklist of the snakes of the Kingdom. The Natural History Journal of Chulalongkorn University 4(1): 47-80 - get paper here
  • David,P. & Vogel, G. 1996. The snakes of Sumatra. An annotated checklist and key with natural history notes. Bücher Kreth, Frankfurt/M.
  • Duméril, A.M.C., G. BIBRON & A.H.A. DUMÉRIL 1854. Erpétologie générale ou Histoire Naturelle complète des Reptiles. Vol. 7 (partie 1). Paris, xvi + 780 S. - get paper here
  • Figueroa A, McKelvy AD, Grismer LL, Bell CD, Lailvaux SP 2016. A Species-Level Phylogeny of Extant Snakes with Description of a New Colubrid Subfamily and Genus. PLoS One 11 (9): e0161070.doi:10.1371/journal.pone.0161070 - get paper here
  • Fukuyama R, Fukuyama I, Kurita T, Kojima Y, Hossman MY, Noda A, Nishikawa K 2021. New herpetofaunal records from Gunung Mulu National Park and its surrounding areas in Borneo. Herpetozoa 34: 89-96 - get paper here
  • Grandison, A. G. C. 1978. Snakes of West Malaysia and Singapore. Annalen des Naturhistorischen Museums in Wien 81 [1977]: 283-303 - get paper here
  • Grandison, A.G.C. 1972. The Gunong Benom Expedition 1967. 5. Reptiles and amphibians of Gunong Benom with a description of a new species of Macrocalamus. Bull. Br. Mus. nat. Hist. (Zool.), London, 23: 45-101. - get paper here
  • GRISMER, L. LEE; EVAN S.H. QUAH, SHAHRUL ANUAR M.S,, MOHD ABDUL MUIN, PERRY L. WOOD, JR & SITI AZIZAH MOHD NOR 2014. A diminutive new species of cave-dwelling Wolf Snake (Colubridae: Lycodon Boie, 1826) from Peninsular Malaysia. Zootaxa 3815 (1): 051–067 - get paper here
  • Grismer, L.L. 2011. Amphibians and reptiles of the Seribuat Archipelago. Edition Chimaira, Frankfurt, 239 pp.
  • Grismer, L.L., McGuire, J.A.; Sosa, R. & Kaiser, H. 2002. Revised checklist and comments on the terrestrial herpetofauna of Pulau Tioman, Peninsular Malaysia. Herpetological Review 33 (1): 26-29 - get paper here
  • Grossmann, W. & Tillack, F. 2004. Pulau Tioman - Perle im Südchinesischen Meer, Teil 1. Reptilia (Münster) 9 (50): 42-49 - get paper here
  • Harrington, Sean M; Jordyn M de Haan, Lindsey Shapiro, Sara Ruane 2018. Habits and characteristics of arboreal snakes worldwide: arboreality constrains body size but does not affect lineage diversification. Biological Journal of the Linnean Society 125 (1): 61–71 - get paper here
  • Hien,P. Grossmann,W. & Schäfer, C. 2001. Beitrag zur Kenntnis der landbewohnenden Reptilienfauna von Pulau Tioman, West-Malaysia. Sauria 23 (4): 11-28 - get paper here
  • Janssen HY, Pham CT, Ngo HT, Le MD, Nguyen TQ, Ziegler T 2019. A new species of Lycodon Boie, 1826 (Serpentes, Colubridae) from northern Vietnam. ZooKeys 875: 1-29 - get paper here
  • Lanza,B. 1999. A new species of Lycodon from the Philippines, with a key to the genus (Reptilia: Serpentes: Colubridae). Tropical Zoology 12: 89-104 - get paper here
  • Lim, K.K.P. & Ng, H.H. 1999. The terrestrial herpetofauna of Pulau Tioman, Peninsular Malaysia. Raffles Bull. Zool., Suppl. No. 6: 131-155 - get paper here
  • Malkmus, R.; Manthey, U.; Vogel, G. Hoffmann, P. & Kosuch, J. 2002. Amphibians and reptiles of Mount Kinabalu (North Borneo). A.R.G. Ganther Verlag, Rugell, 404 pp.
  • Manthey, U. & Grossmann, W. 1997. Amphibien & Reptilien Südostasiens. Natur und Tier Verlag (Münster), 512 pp. - get paper here
  • Manthey,U. 1983. Exkursion am Mt. Kinabalu (4101 m), Nordborneo, Teil 3: Checkliste der Herpetofauna oberhalb 600 m ü. NN. Herpetofauna 5 (23): 20-31 - get paper here
  • Murphy, J. C.; Voris, H. K.; Karns, D. R. 1994. A field guide and key to the snakes of the Danum Valley, A bornean tropical rainforest ecosystem. Bull. Chicago Herp. Soc. 29: 133-151
  • Onn, Chan Kin; Mohd. Shahfiz Azman, Nor Azlin and Pan Khang Aun 2009. Additions to the Herpetofauna of Pasoh Forest Reserve, Negeri Sembilan, Peninsular Malaysia. Tropical Life Sciences Research, 20 (1): 71–80 - get paper here
  • Rooijen, J. van & van Rooijen, M. 2002. Einige Ergänzungen, Berichtigungen und neue Beobachtungen zur Herpetofauna von Pulau Tioman, West-Malaysia. Sauria 24 (3): 3-12 [erratum in 24 (4): 34] - get paper here
  • Stoliczka,F. 1873. Notes on some species of Malayan Amphibia and reptilia. J. Asiat. Soc. Bengal 42: 111-126 - get paper here
  • Stuebing, R.B. & INGER, R.F. 1999. A field guide to the snakes of Borneo. Natural history Publications (Borneo), Kota Kinabalu, 254 pp. [corrections in HR 31: 124]. - get paper here
  • Stuebing, R.B., Inger, R.F. & Lardner, B. 2014. A field guide to the snakes of Borneo, second edition. Natural history Publications (Borneo)
  • Syafiq MF, Badli-Sham BH, Grismer LL, Ahmad AB 2023. Uneven species occurrence and richness of lowland snakes (Serpentes, Squamata) in Terengganu, Peninsular Malaysia, with new locality records. ZooKeys 1168: 11–39
  • Syafiq, Muhamad Fatihah; Baizul Hafsyam Badli-Sham, Noor-Shahirah Ibrahim, Lukman H. B. Ismail, Muhammad Afif Mohd Amin, Goh Shu Xian, Rabiatul ‘Aqilah Mohamad Ariffin, Muhammad Afiq-Suhaimi, Lok Kar Men, Dila Adilina Danelo, Mohamad Aqmal-Naser, Muh 2024. TAXONOMIC COMPOSITION, DIVERSITY, AND CONSERVATION STATUS OF REPTILIAN FAUNA AT SEKAYU LOWLAND FORESTS, TERENGGANU, PENINSULAR MALAYSIA. Russian Journal of Herpetology Vol. 31, No. 1, 2024, pp. 14 – 23 - get paper here
  • TEYNIÉ, ALEXANDRE; PATRICK DAVID, & ANNEMARIE OHLER 2010. Note on a collection of Amphibians and Reptiles from Western Sumatra (Indonesia), with the description of a new species of the genus Bufo. Zootaxa 2416: 1–43 - get paper here
  • Vogel, G. & R. Kreutz 1990. Erstnachweis von Lepturophis borneensis BOULENGER 1900 für Thailand. Sauria 12 (1): 21-23 - get paper here
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp.
  • WOSTL, ELIJAH; AMIR HAMIDY, NIA KURNIAWAN, ERIC N. SMITH 2017. A new species of Wolf Snake of the genus Lycodon H. Boie in Fitzinger (Squamata: Colubridae) from the Aceh Province of northern Sumatra, Indonesia. Zootaxa 4276 (4): 539–553 - get paper here
 
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