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Lycodon futsingensis (POPE, 1928)

IUCN Red List - Lycodon futsingensis - Least Concern, LC
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Higher TaxaColubridae, Colubrinae, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies 
Common NamesChinese: 福清白环蛇 
SynonymDinodon futsingensis POPE 1928
Dinodon septentrionale ruhstrati — MAKI 1931 (part.)
Dinodon futsingensis — BOURRET 1934
Dinodon futsingensis — BOURRET 1935: 6
Dinodon futsingense — BOURRET 1936
Lycodon ruhstrati — POPE 1935 (part.)
Lycodon ruhstrati ruhstrati — OTA 1988 (part.)
Lycodon ruhstrati ruhstrati — LANZA 1999 (part.)
Ophites ruhstrati — ZHAO & ADLER 1993 (part.)
Lycodon futsingensis — VOGEL et al. 2009
Lycodon futsingensis — GUO et al. 2013
Lycodon futsingensis — WALLACH et al. 2014: 394
Lycodon futsingensis — WANG et al. 2022 
DistributionChina (Fukien = Fujian, Hong Kong, Guangdong)
Vietnam (Hoa Binh, Vin Puh, Bắc Kạn [Ngan-Son], Lào Cai, Hà Tĩnh, Quang Binh [Phong Nha-Ke Bang National Park], Ha Giang), Laos

Type locality: “Futsing Hsien, Fukien Province”, now Fuqing Xian [County], 25°44’N, 119°22’E, Fujian Province, People’s Republic of China.  
Reproductionoviparous. 
TypesHolotype: AMNH 34106, collected by C.H. Pope, August-October, 1925. – Paratype. AMNH 34105, same data as the holotype. 
DiagnosisDiagnosis: A species of the genus Lycodon, characterized by: (1) a loreal not entering orbit; (2) 17 dorsal scale rows at the forepart of the body and at midbody; (3) the upper dorsal and vertebral scale rows entirely smooth; (4) 193-204 ventrals in males and 198-208 in females; (5) 72-87 SC in males and 78-85 in females; (6) a relative tail length ratio of 0.194-0.229 in males and 0.194-0.216 in females; (7) 19-33 light bands on a dark body; (8) the first band starting at Ve 13-21; and (9) upper maxillary teeth: 7-8 + 3-4 + 2-3. This species can be recognized by the combination of unkeeled dorsal scale rows and low ventral and subcaudal counts. For a detailed comparisons with other species of Lycodon see discussion in VOGEL et al. (2009). 
CommentSynonymy: the species has been synonymized with Lycodon ruhstrati (Pope, 1929), to which it is morphologically similar (Pope, 1935; Vogel et al., 2009). This arrangement has been followed for a long time by some authors (Zhao et al., 1998; Lanza, 1999). Recently, on the basis of morphological comparison, Vogel et al. (2009) revalidated Dinodon futsingensis and recognized it as Lycodon futsingensis. 
EtymologyThis species was named according to the name of the type locality, Futsing Hsien, now spelt as Fuxing Qian in Pinyin. 
References
  • Bourret, R. 1934. Notes herpétologiques sur l'Indochine Française II. Sur quelques serpents des montagnes du Tonkin. Bull. Gen. Instr. Pub. Hanoi 8 (April 1934): 149-157 [13: 1-11 ?]
  • Bourret, R. 1936. Les Serpents de l'Indochine, vol. 2. Imprimerie Henri Basuyau & Cie, Toulouse.
  • Bourret,R. 1935. Notes herpétologiques sur l'Indochine Française X. Les serpents de la station d’altitude du Tam-dao. Bull. Gen. Instr. Pub. Hanoi 8 (avril): 259-271 (1-13)
  • Deuve, J. 1961. Liste annotee des Serpents du Laos. Bull. Soc. Sci. Nat. Laos 1:5-32.
  • Francis, A. 2021. A field guide to the snakes of Hong Kong. Regal Printing, Hong Kong - get paper here
  • Guo, Peng; Liang Zhang, Qin Liu, Cao Li, R. Alexander Pyron, Ke Jiang, Frank T. Burbrink 2013. Lycodon and Dinodon: One genus or two? Evidence from molecular phylogenetics and morphological comparisons. Molecular Phylogenetics and Evolution 68 (1): 144–149 - get paper here
  • Hecht, Vera L.; Cuong T. Pham, Tao T. Nguyen, Truong Q. Nguyen, Michael Bonkowski & Thomas Ziegler 2013. First report on the herpetofauna of Tay Yen Tu Nature Reserve, northeastern Vietnam. Biodiversity Journal 4 (4): 507–552
  • Janssen HY, Pham CT, Ngo HT, Le MD, Nguyen TQ, Ziegler T 2019. A new species of Lycodon Boie, 1826 (Serpentes, Colubridae) from northern Vietnam. ZooKeys 875: 1-29 - get paper here
  • Lanza,B. 1999. A new species of Lycodon from the Philippines, with a key to the genus (Reptilia: Serpentes: Colubridae). Tropical Zoology 12: 89-104 - get paper here
  • Luu, V. Q., Dinh, T. S., Lo, O. V., Nguyen, T. Q. and Ziegler, T. 2020. New records and an updated list of reptiles from Ba Vi National Park, Vietnam. Bonn Zoological Bulletin 69: 1-9.
  • Luu, Vinh Quang; Michael Bonkowski, Truong Quang Nguyen, Minh Duc Le,, Thomas Calame & Thomas Ziegler 2018. A new species of Lycodon Boie, 1826 (Serpentes: Colubridae) from central Laos. Revue suisse de Zoologie 125(2): 263-276
  • LUU, VINH QUANG; THOMAS ZIEGLER, NGHIA VAN HA, MINH DUC LE, TUOI THI HOANG 2019. A new species of Lycodon Boie, 1826 (Serpentes: Colubridae) from Thanh Hoa Province, Vietnam. Zootaxa 4586 (2): 261-277 - get paper here
  • Maki, M. 1931. Monograph of the Snakes of Japan. Dai-ichi Shobo, Tokyo. (1), 7, 240 pages (expanded edition 1933)
  • NEANG, THY; TIMO HARTMANN, SEIHA HUN, NICHOLAS J. SOUTER & NEIL M. FUREY 2014. A new species of wolf snake (Colubridae: Lycodon Fitzinger, 1826) from Phnom Samkos Wildlife Sanctuary, Cardamom Mountains, southwest Cambodia. Zootaxa 3814 (1): 068–080 - get paper here
  • Nguyen TV, Brakels P, Maury N, Sudavanh S, Pawangkhanant P, Idiiatullina S, Lorphengsy S, Inkhavilay K, Suwannapoom C, Poyarkov NA. 2020. New herpetofaunal observations from Laos based on photo records. Amphibian & Reptile Conservation 14(2) [General Section]: 218–249 (e248) - get paper here
  • Nguyen, Luan Thanh; Ha Van Hoang, Thang Tai Nguyen, McCormack Timothy E.M., Sang Ngoc Nguyen 2016. A COLLECTION OF AMPHIBIANS AND REPTILES FROM BAC HUONG HOA NATURE RESERVE, QUANG TRI PROVINCE, VIETNAM. Proceedings of the 3nd National Scientific Workshop “Amphibians and Reptiles in Vietnam - get paper here
  • Nguyen, Truong Quang 2017. New records of amphibians and reptiles from Ha Giang Province, Vietnam. Herpetology Notes 10: 183-191 - get paper here
  • Nguyen, Truong Quang; Tan Van Nguyen, Cuong The Pham, An Vinh Ong & Thomas Ziegler 2018. New records of snakes (Squamata: Serpentes) from Hoa Binh Province, northwestern Vietnam. Bonn zoological Bulletin 67 (1): 15–24 - get paper here
  • Ota, Hidetoshi 1988. Taxonomic notes on Lycodon ruhstrati (Colubridae: Ophidia) from East Asia. Journal of Taiwan Museum 41 (1): 85-91
  • Pham AV, Ziegler T, Nguyen TQ 2020. New records and an updated checklist of snakes from Son La Province, Vietnam. Biodiversity Data Journal 8: e52779 - get paper here
  • Pope, CLIFFORD H. 1928. Seven new reptiles from Fukien Province, China. American Museum Novitates 320: 1-6. - get paper here
  • Pope, CLIFFORD H. 1935. The Reptiles of China. Turtes, Crocodilians, Snakes, Lizards. Amer. Mus. Nat. Hist., New York, Nat. Hist. Central Asia, 10: lii, 1-604 - get paper here
  • Vogel, G. & Luo, J. 2011. A new species of the genus Lycodon (Boie, 1826) from the southwestern mountains of China (Squamata: Colubridae). Zootaxa 2807: 29–40 - get paper here
  • Vogel, G., T. Q. Nguyen, P. Kingsada & T. Ziegler 2012. A new species of the genus Lycodon BOIE, 1826 from Laos (Squamata: Colubridae). North-Western Journal of Zoology, 8: 344-352 - get paper here
  • Vogel, G.; David, P.; Pauwels, O.S.G.; Sumontha, M.; Norval, G.; Hendrix, R.; Vu, N.T. & Ziegler, T. 2009. A revision of Lycodon ruhstrati (Fischer 1886) auctorum (Squamata Colubridae), with the description of a new species from Thailand and a new subspecies from the Asian mainland. Tropical Zoology 22: 131-182 - get paper here
  • Vogel, Gernot 2010. Zur Systematik der Schlangen der orientalischen Region: Forschungsdefizite führen zur Unterschätzung der Artenvielfalt. Ophidia 4 (2): 18-26 - get paper here
  • Vogel, Gernot & Patrick David 2010. A new species of the genus Lycodon (Boie, 1826) from Yunnan Province, China (Serpentes: Colubridae). Bonn Zool. Bull. 57 (2): 289–296 - get paper here
  • VOGEL, GERNOT; PATRICK DAVID 2019. A new species of the Lycodon fasciatus complex from the Khorat Plateau, eastern Thailand (Reptiles, Squamata, Colubridae). Zootaxa 4577 (3): 515–528 - get paper here
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp.
  • Wang, C. G. M. Z. X., Jia, S. Y. K. T. L., & Hu, C. Z. J. 2022. An updated species checklist of amphibians and reptiles in Fujian Province, China. Biodiversity Science, 22090 - get paper here
  • Wang, Kai; Jinlong Ren, Hongman Chen, Zhitong Lyu, Xianguang Guo Ke Jiang, Jinmin Chen, Jiatang Li, Peng Guo, Yingyong Wang, Jing Che 2020. The updated checklists of amphibians and reptiles of China. Biodiversity Science 28 (2): 189-218 - get paper here
  • ZHANG Liang; JIANG Ke, HU Ping, YU Bao-Cheng, PENG Bo-Yong, TANG Xiao-Ping, HU Hui-Jian 2011. Lycodon futsingensis: A New Snake Record in Guangdong Province,China. Chinese Journal of Zoology 46 (1):128-130
  • Zhang, Liang and Kai Wang. 2014. Lycodon futsingensis (Fuqing wolf snake) diet. Herpetological Review 45 (4): 711-712 - get paper here
  • Zhao,E. & Adler,K. 1993. Herpetology of China. SSAR, Oxford/Ohio, 1-522
 
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