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Lycodon davidi VOGEL, NGUYEN, KINGSADA & ZIEGLER, 2012

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Higher TaxaColubridae, Colubrinae, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Common NamesE: David’s Wolf Snake
G: Davids Wolfszahnnatter
Vietnamese: Rắn khuyết da-vid 
SynonymLycodon davidi VOGEL, NGUYEN, KINGSADA & ZIEGLER 2012
Lycodon davidi — WALLACH et al. 2014: 393 
DistributionLaos (Vientiane)

Type locality: Nam Cave 268 m elevation, near Ban Muang Song, Vang Vieng District, Vientiane Province, Laos  
TypesHolotype: IEBR A.2011.7, male (ventral tail base dissected), (Figs 1–3, 5, 6), collected by Truong Quang Nguyen and Phouthone Kingsada on 6 May 2010. 
DiagnosisDiagnosis. A species of the genus Lycodon characterized by: (1) loreal scale not entering orbit; (2) 17 dorsal scale rows at the forepart of the body and 17 dorsal scale rows at midbody; (3) dorsal scale rows keeled; (4) 224 ventrals in the single known male, females unknown; (5) 99 SC in the single known male; (6) a relative tail length of about 0.209 in the single known male (7) 8 supralabials with SL 3–5 touching the orbit; (7) 84 narrow brownish-grey bands on a dark brown body, widening laterally; (8) width of the first band about 1.0 scale on the vertebral row, about 3 ventrals at its base; and (9) the venter heavily dark speckled in the posterior two thirds.

The new species can be recognized by the combination of the loreal scale not entering orbit, the high number of narrow dorsal bands and especially the high number of bands on the tail and the venter which is heavily speckled in the posterior two thirds. Detailed comparisons with other species of the genus Lycodon are provided in the Discussion. 
CommentHabitat: evergreen karst forest, possibly in caves as the type specimen was found at the entrance of a karst cave. 
EtymologyThis species is named in honour of our friend and colleague Patrick David (MNHN, Paris, France), for his outstanding contributions to the reptile fauna of the Oriental Region. 
  • Janssen HY, Pham CT, Ngo HT, Le MD, Nguyen TQ, Ziegler T 2019. A new species of Lycodon Boie, 1826 (Serpentes, Colubridae) from northern Vietnam. ZooKeys 875: 1-29 - get paper here
  • Luu, Vinh Quang; Michael Bonkowski, Truong Quang Nguyen, Minh Duc Le,, Thomas Calame & Thomas Ziegler 2018. A new species of Lycodon Boie, 1826 (Serpentes: Colubridae) from central Laos. Revue suisse de Zoologie 125(2): 263-276
  • NEANG, THY; TIMO HARTMANN, SEIHA HUN, NICHOLAS J. SOUTER & NEIL M. FUREY 2014. A new species of wolf snake (Colubridae: Lycodon Fitzinger, 1826) from Phnom Samkos Wildlife Sanctuary, Cardamom Mountains, southwest Cambodia. Zootaxa 3814 (1): 068–080 - get paper here
  • Vogel, G., T. Q. Nguyen, P. Kingsada & T. Ziegler 2012. A new species of the genus Lycodon BOIE, 1826 from Laos (Squamata: Colubridae). North-Western Journal of Zoology, 8: 344-352 - get paper here
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp.
  • Ziegler, T & Nguyen, QT 2015. Neues von den Forschungs- und Naturschutzprojekten in Vietnam und Laos. Zeitschrift des Kölner Zoos 2015 (2): 79-
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