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Lycodon ruhstrati (FISCHER, 1886)

IUCN Red List - Lycodon ruhstrati - Least Concern, LC

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Higher TaxaColubridae, Colubrinae, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
SubspeciesLycodon ruhstrati abditus VOGEL, DAVID, PAUWELS, SUMONTHA, NORVAL, HENDRIX, RU & ZIEGLER 2009
Lycodon ruhstrati ruhstrati (FISCHER 1886) 
Common NamesE: Mountain Wolf Snake, Formosa Wolf Snake
G: Berg-Wolfszahnnatter
Chinese: 黑背白环蛇 
SynonymOphites Ruhstrati FISCHER 1886: 16
Dinodon septentrionalis — BOULENGER 1893: 363 (part.)
Dinodon septentrionalis var. ruhstrati — BOULENGER 1899: 165
Dinodon septentrionale ruhstrati — STEJNEGER 1907: 370
Ophites ruhstrati — ZHAO & ADLER 1993
Lycodon ruhstrati — LANZA 1999
Dinodon ruhstrati — WEB SITE
Lycodon ruhstrati — ZIEGLER 2002: 241
Lycodon cf. ruhstrati — ZIEGLER et al. 2007
Lycodon ruhstrati — WALLACH et al. 2014: 395
Lycodon ruhstrati — WANG et al. 2022

Lycodon ruhstrati abditus — LUU et al. 2018 
DistributionTaiwan, China (Jiangxi, Fujian, Guangdong, northward to Anhui and west to Sichuan [elevation 800-1850 m] and SE Gansu), N Vietnam (elevation 500-1500 m)

ruhstrati: Taiwan (endemic)

abditus: Vietnam, Laos. Type locality: U Bo region, Phong Nha – Ke Bang NP, Quang Binh Province, Vietnam.

Type locality: Süd-Formosa [= South Taiwan], China.  
TypesSyntypes: LMNMO 918 (formerly LMNMO 37c & 73), a 583+ mm male, and LMNMO 919 (formerly LMNMO 37b & 74)
Syntypes. REP 918-19
Holotype: ZFMK 86451 (GenBank: EU999209), adult female. Collected by Thomas Ziegler, 20 June 2006 (mentioned as Lycodon cf. ruhstrati in Ziegler et al. 2007), paratypes in ZFMK, MNHN and ZMB [abditus] 
DiagnosisDiagnosis. A species of the genus Lycodon, is characterized by: (1) a loreal not entering the orbit; (2) 17 dorsal scale rows at the forepart of the body and at midbody; (3) upper dorsal rows distinctly keeled; and (4) a banded body and tail.

Diagnosis (ruhstrati): A species of the genus Lycodon, characterized by: (1) a loreal not entering orbit; (2) 17 dorsal scale rows at the forepart of the body and at midbody; (3) upper dorsal rows distinctly keeled; (4) 212-228 ventrals in males and 217-224 in females; (5) 105-114 Sc in males and 97-108 in females; (6) a relative tail length of 0.220-0.248 in males and 0.211-0.239 in females; (7) 33-46 bands on body; (8) the first band starting at Ven 8-16.

Diagnosis (abditus): A subspecies of Lycodon ruhstrati, characterized by: (1) a loreal not entering orbit; (2) 17 dorsal scale rows at the forepart of the body and at midbody; (3) three to five upper dorsal scale (including vertebral) rows distinctly keeled; (4) 214-224 ventrals in males and 220-229 in females; (5) 90-100 Sc in males and 92-103 in females; (6) a relative tail length of 0.208-0.237 in males and 0.207-0.236 in females; (7) 19-43 bands on body; (8) the first band starting at Ve 12-17; and (9) 6-7 + 3-4 + 2 much enlarged upper maxillary teeth, each group of teeth separated by a wide gap.
This new subspecies differs from the nominate subspecies by (1) a lower ratio of TaL/TL in males, 0.208-0.237 (x = 0.218, s = 0.009) vs 0.220-0.248 (x = 0.230, s = 0.009) in L. ruhstrati ruhstrati, (2) a lower number of SC, 90-100 (x = 93.5, s = 3.2) in males and 92-103 (x = 96.0, s = 4.1) in females vs 105-114 (x = 107.8, s = 3.4) and 97-106 (x = 103.1, s = 3.8) respectively in the nominate subspecies, and (3) a lower number of bands on the body, 29-43 (x = 34.6, s = 3.6) vs 33-46 (x = 38.6, s = 3.6) in the nominate subspecies. 
CommentTaxonomy and nomenclature: For a discussion of nomenclatorial issues concerning the genera Lycodon and Ophites see Zhao & Adler (who listed this species under Ophites). Dinodon futsingensis POPE 1928, previously a synonym of L. ruhstrati, has been revalidated by VOGEL et al. 2009. Lycodon ruhstrati multifasciatus (MAKI 1931) has been elevated to full species status by VOGEL et al. 2009.

Habitat: partly arboreal (Harrington et al. 2018). 
EtymologyNamed after Mr. Ruhstrat, who collected the types in southern Taiwan.

abditus: The subspecific nomen of this subspecies is the Latin adjective abditus (-a, -um), meaning “hidden”, in allusion to the long confusion of this species with Lycodon ruhstrati ruhstrati. 
  • Böhme, Wolfgang 2014. Herpetology in Bonn. Mertensiella 21. vi + 256 pp. - get paper here
  • Boulenger, G.A. 1893. Catalogue of the snakes in the British Museum (Nat. Hist.) I. London (Taylor & Francis), 448 pp. - get paper here
  • Boulenger, GEORGE A. 1899. On a collection of reptiles and batrachians made by Mr. J. D. La Touche in N.W. Fokien, China. Proc. Zool. Soc. London 1899: 159-172 - get paper here
  • Fischer, J. G. 1886. Herpetologische Notizen. Abh. Geb. Naturw. Hamb. 9: 51-67 (1-19) - get paper here
  • Harrington, Sean M; Jordyn M de Haan, Lindsey Shapiro, Sara Ruane 2018. Habits and characteristics of arboreal snakes worldwide: arboreality constrains body size but does not affect lineage diversification. Biological Journal of the Linnean Society 125 (1): 61–71 - get paper here
  • Janssen HY, Pham CT, Ngo HT, Le MD, Nguyen TQ, Ziegler T 2019. A new species of Lycodon Boie, 1826 (Serpentes, Colubridae) from northern Vietnam. ZooKeys 875: 1-29 - get paper here
  • KUCHARZEWSKI, CH. & TILLACK, F. & FUHRMANN, K. 2009. Rediscovery and redescription of the type specimens of Ophites ruhstrati FISCHER, 1886. Herpetozoa 22: - get paper here
  • Lanza,B. 1999. A new species of Lycodon from the Philippines, with a key to the genus (Reptilia: Serpentes: Colubridae). Tropical Zoology 12: 89-104 - get paper here
  • Luu, V. Q., Dinh, T. S., Lo, O. V., Nguyen, T. Q. and Ziegler, T. 2020. New records and an updated list of reptiles from Ba Vi National Park, Vietnam. Bonn Zoological Bulletin 69: 1-9.
  • Luu, Vinh Quang; Michael Bonkowski, Truong Quang Nguyen, Minh Duc Le,, Thomas Calame & Thomas Ziegler 2018. A new species of Lycodon Boie, 1826 (Serpentes: Colubridae) from central Laos. Revue suisse de Zoologie 125(2): 263-276
  • Mori,M. 1982. Japans Schlangen, Vols. 1-3 (80, 102, 123 pp.). Tokyo (Igaku-Shoin Ltd.) [bilingual edition in German and Japanese]
  • NEANG, THY; TIMO HARTMANN, SEIHA HUN, NICHOLAS J. SOUTER & NEIL M. FUREY 2014. A new species of wolf snake (Colubridae: Lycodon Fitzinger, 1826) from Phnom Samkos Wildlife Sanctuary, Cardamom Mountains, southwest Cambodia. Zootaxa 3814 (1): 068–080 - get paper here
  • Nguyen, S.V., Ho, C.T. and Nguyen, T.Q. 2009. Herpetofauna of Vietnam. Chimaira, Frankfurt, 768 pp.
  • NORVAL, G. & J.-J. MAO 2006. Die Berg-Wolfszahnnatter (Lycodon ruhstrati ruhstrati [FISCHER, 1886]): Eine Beschreibung ihres Abwehrverhaltens. Sauria 28 (4): 47-50 - get paper here
  • Norval, G. & Mao, J.J. 2008. An instance of arboricolous predation by a Mountain Wolf Snake (Lycodon ruhstrati ruhstrati [FISCHER], 1886]) on a Brown anole (Norops sagrei DUMÉRIL & BIBRON, 1837). Sauria 30 (1): 59-62 - get paper here
  • Norval, G. & Mao, J.J. 2008. An instance of Japalura swinhonis as prey of Lycodon ruhstrati ruhstrati in Chiayi County, Taiwan. Sauria 30 (3): 43-45 - get paper here
  • Norval, Gerrut; Charles R. Bursey, Stephen R. Goldberg and Jean-Jay Mao. 2009. The first record of Kiricephalus pattoni Stephens, 1908 as a parasite of Lycodon ruhstrati ruhstrati Fischer, 1886 from Chiayi County, Taiwan. Herpetology Notes 2: 79-80. - get paper here
  • Orlov, N. L., R. W. Murphy & T. J. Papenfuss 2000. List of snakes of Tam-Dao mountain ridge (Tonkin, Vietnam). Russ. J. Herpetol. 7 (1): 69-80. - get paper here
  • Ota, H. 2000. Current status of the threatened amphibians and reptiles of Japan. Popul. Ecol. 42: 5-9 - get paper here
  • Ota, Hidetoshi 1988. Taxonomic notes on Lycodon ruhstrati (Colubridae: Ophidia) from East Asia. Journal of Taiwan Museum 41 (1): 85-91
  • Pope, CLIFFORD H. 1928. Seven new reptiles from Fukien Province, China. American Museum Novitates 320: 1-6. - get paper here
  • Stejneger, L. 1910. The batrachians and reptiles of Formosa. Proc. US Natl. Mus. 38: 91-114 - get paper here
  • Stejneger, LEONHARD H. 1907. Herpetology of Japan and adjacent territory. Bull. US Natl. Mus. 58: xx, 1-577 - get paper here
  • Vogel, G. & Brachtel, N. 2008. Contribution to the knowledge of Lycodon ruhstrati (FISCHER, 1886) in Vietnam - taxonomy and biology of a little-known species. Salamandra 44 (4): 207-224 - get paper here
  • Vogel, G. & Luo, J. 2011. A new species of the genus Lycodon (Boie, 1826) from the southwestern mountains of China (Squamata: Colubridae). Zootaxa 2807: 29–40 - get paper here
  • Vogel, G., T. Q. Nguyen, P. Kingsada & T. Ziegler 2012. A new species of the genus Lycodon BOIE, 1826 from Laos (Squamata: Colubridae). North-Western Journal of Zoology, 8: 344-352 - get paper here
  • Vogel, G.; David, P.; Pauwels, O.S.G.; Sumontha, M.; Norval, G.; Hendrix, R.; Vu, N.T. & Ziegler, T. 2009. A revision of Lycodon ruhstrati (Fischer 1886) auctorum (Squamata Colubridae), with the description of a new species from Thailand and a new subspecies from the Asian mainland. Tropical Zoology 22: 131-182 - get paper here
  • Vogel, Gernot 2010. Zur Systematik der Schlangen der orientalischen Region: Forschungsdefizite führen zur Unterschätzung der Artenvielfalt. Ophidia 4 (2): 18-26 - get paper here
  • Vogel, Gernot & Patrick David 2010. A new species of the genus Lycodon (Boie, 1826) from Yunnan Province, China (Serpentes: Colubridae). Bonn Zool. Bull. 57 (2): 289–296 - get paper here
  • VOGEL, GERNOT; PATRICK DAVID 2019. A new species of the Lycodon fasciatus complex from the Khorat Plateau, eastern Thailand (Reptiles, Squamata, Colubridae). Zootaxa 4577 (3): 515–528 - get paper here
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp.
  • Wang, C. G. M. Z. X., Jia, S. Y. K. T. L., & Hu, C. Z. J. 2022. An updated species checklist of amphibians and reptiles in Fujian Province, China. Biodiversity Science, 22090 - get paper here
  • Wang, Kai; Jinlong Ren, Hongman Chen, Zhitong Lyu, Xianguang Guo Ke Jiang, Jinmin Chen, Jiatang Li, Peng Guo, Yingyong Wang, Jing Che 2020. The updated checklists of amphibians and reptiles of China. Biodiversity Science 28 (2): 189-218 - get paper here
  • Zhao, E. (ed.) 2003. Coloured atlas of Sichuan reptiles [in Chinese]. Beijing, China Forestry Publishing House
  • Zhao, E.M. 2006. The snakes of China [in Chinese]. Hefei, China, Anhui Sience & Technology Publ. House, Vol. I, 372 pp., Vol. II (color plates), 280 pp.
  • Zhao,E. & Adler,K. 1993. Herpetology of China. SSAR, Oxford/Ohio, 1-522
  • Ziegler, T. 2002. Die Amphibien und Reptilien eines Tieflandfeuchtwald-Schutzgebietes in Vietnam. Natur und Tier Verlag (Münster), 342 pp. - get paper here
  • ZIEGLER, THOMAS; RALF HENDRIX, VU NGOC THANH, MARTINA VOGT, BERNHARD FORSTER & DANG NGOC KIEN 2007. The diversity of a snake community in a karst forest ecosystem in the central Truong Son, Vietnam, with an identification key. Zootaxa 1493: 1-40 - get paper here
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