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Lygodactylus winki VENCES, MULTZSCH, GIPPNER, MIRALLES, CROTTINI, GEHRING, RAKOTOARISON, RATSOAVINA, GLAW & SCHERZ, 2022

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common Names 
SynonymLygodactylus winki VENCES, MULTZSCH, GIPPNER, MIRALLES, CROTTINI, GEHRING, RAKOTOARISON, RATSOAVINA, GLAW & SCHERZ 2022 
DistributionMadagascar (Ampotsidy, Andrevorevo, Bemanevika)

Type locality: Ampotsidy, 15.7 km NNW of Bealanana (8.7 km NNW of Beandrarezona), northern Madagascar, S 14.41900, E 48.71883, 1364 m a.s.l.  
Reproduction 
TypesHolotype. ZSM 47/2016 (MSZC 0075), adult male, collected by M.D. Scherz, J. Borrell, L. Ball, T. Starnes, E. Razafimandimby, D.H. Nomenjanahary, and J. Rabearivony, on 22 December 2015.
Paratypes. ZSM 48/2016 (MSZC 0110), adult male, collected by same collectors and at same locality as holotype, at geographical coordinates S14.42843, E48.72285, 1315 m a.s.l., on 29 December 2015; UADBA-R 70856–70859 (MSZC 0011, 0019, 0023, 0077), two males and two females, respectively, collected by same collectors and at same locality as holotype, between coordinates S14.41455–14.42317, E48.71149–48.71916, 1320–1404 m
a.s.l., on 18–22 December 2015; ZSM 1763/2010 (ZCMV 12502), by M. Vences, D.R. Vieites, R.D. Randrianiaina, F.M. Ratsoavina, S. Rasamison, A. Rakotoarison, E. Rajeriarison, and T. Rajoafiarison, at Bemanevika, Antsirakala campsite, geographical coordinates S14.43061, E48.60179, 1466 m a.s.l., on 27 June 2010; ZSM 555/2014 (DRV 6288), collected by F.M. Ratsoavina, D.R. Vieites, M. Vences, R.D. Randrianiaina, S. Rasamison, A. Rakotoarison, E. Rajeriarison, and T. Rajoafiarison at Andrevorevo, geographical coordinates S14.3464, E49.1028, 1717 m a.s.l., on 21 June 2010. 
DiagnosisDiagnosis: “A lineage forming part of the subclade A5 of the subgenus Domerguella. From the other species in this subclade it differs as follows:
- from L. miops by a lower longitudinal count of ventral scales despite minimal overlap (83–98 vs. 98–113)
- from L. guibei by apparently relatively longer hindlimbs (HIL/SVL 0.49–0.54 vs. 0.42–0.49), more expressed supraocular bulges, and proportionally longer and higher heads with non-overlapping values (relative snout tip to tympanum distance in percent, 24.7–28.8 vs. 23.0–24.6; relative head height in percent, 12.5–14.2 vs. 11.1–12.4).
- from L. fritzi by more dorsolateral tubercles between limbs (5–8 vs. 2), lower longitudinal dorsal scale count (187–222 vs. 247), and weakly expressed lateral tubercles at tail base vs. clearly recognizable usually large in males and females
- from L. ulli by a lower longitudinal counts of dorsal scales and ventral scales.” 
CommentNatural history: The holotype was collected in primary rainforest, on the trunk of a big tree, 0.2 m above the ground. The paratypes from Ampotsidy were mostly collected at night sleeping on leaves, twigs, or vines. 
EtymologyDedicated to Michael Wink, pharmacologist, herpetologist, ornithologist and professor emeritus of the University of Heidelberg, in recognition for his support of research in squamate systematics. 
References
  • VENCES, M., MULTZSCH, M., GIPPNER, S., MIRALLES, A., CROTTINI, A., GEHRING, P. S., ... & SCHERZ, M. D. 2022. Integrative revision of the Lygodactylus madagascariensis group reveals an unexpected diversity of little brown geckos in Madagascar’s rainforest. Zootaxa 5179 (1): 1-61 - get paper here
 
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