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Lygosoma isodactylum (GÜNTHER, 1864)

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Higher TaxaScincidae, Lygosominae, Scincoidea, Sauria, Squamata (lizards)
Common NamesE: Siamese Writhing Skink, Even-toed Supple Skink
G: Gleichfinger-Laubskink 
SynonymEumeces isodactylus GÜNTHER 1864: 93
Lygosoma isodactylum — BOULENGER 1887: 339
Riopa isodactyla — SMITH 1935: 317
Squamicilia isodactyla — MITTLEMAN 1952
Riopa isodactyla — TAYLOR 1963: 968
Lygosoma isodactyla (Riopa isodactyla) — COX et al. 1998: 114
Riopa isodactyla — CHAN-ARD et al. 1999: 27
Lygosoma isodactylum — COTA et al. 2011
Lygosoma isodactylum — FREITAS et al. 2019 
DistributionCambodia, C Thailand (between Ayuthia and Paknam Po [Nakon Sawan])  
TypesHolotype: BMNH 1946.8.10.69 (formerly, Camboja, collected M. Mouhot. 
DiagnosisDiagnosis. According to Günther (1864), Boulenger (1887), Taylor (1963) and Bourret (2009) the species shows the following diagnostic features: SVL 83–117 mm, longer than TaL; lower eyelids scaly; supranasals separated by rostral and fused with nasal anteriorly; frontal little longer than frontoparietal; frontoparietal single; nuchals absent; ear openings almost covered by two large scales; supralabials 7; scale rows around midbody 30–34; paravertebral scales 88–98; limbs very short (SVL/FlL: about 9; SVL/HlL: about 7); subdigital lamellae under fourth toe 7–10 (see Tables 1, 2) (Geissler et al. 2011). 
CommentType species: Eumeces isodactylus is the type species of the genus Squamicilia.

MITTLEMAN (1952) defined the genus Squamicilia (type species: Eumeces isodactylus) as follows: “Differs essentially from Riopa as follows: Lower eyelid scaly, lacking a more or less transparent or translucent disc; supranasals reduced and not forming a median suture; frontoparietals usually united, interparietal much reduced; ear opening reduced, often only a small oblique slit”. 
  • Boulenger, G. A. 1887. Catalogue of the lizards in the British Museum (Nat. Hist.) III. Lacertidae, Gerrhosauridae, Scincidae, Anelytropsidae, Dibamidae, Chamaeleontidae. London: 575 pp. - get paper here
  • Chan-ard, T., Parr, J.W.K. & Nabhitabhata, J. 2015. A field guide to the reptiles of Thailand. Oxford University Press, NY, 352 pp. [see book reviews by Pauwels & Grismer 2015 and Hikida 2015 for corrections] - get paper here
  • Cota, Michael; Peter Geissler , Tanya Chan-ard and Sunchai Makchai 2011. First Record of Lygosoma angeli (Smith, 1937) (Reptilia: Squamata: Scincidae) in Thailand with Notes on Other Specimens from Laos. The Thailand Natural History Museum Journal 5(2): 125-132
  • Cox, Merel J.; Van Dijk, Peter Paul; Jarujin Nabhitabhata & Thirakhupt,Kumthorn 1998. A Photographic Guide to Snakes and Other Reptiles of Peninsular Malaysia, Singapore and Thailand. Ralph Curtis Publishing, 144 pp.
  • FREITAS, ELYSE S.; ANIRUDDHA DATTA-ROY, PRAVEEN KARANTH, L. LEE GRISMER and CAMERON D. SILER 2019. Multilocus phylogeny and a new classification for African, Asian and Indian supple and writhing skinks (Scincidae: Lygosominae). Zoological Journal of the Linnean Society 186 (4): 1067–1096 - get paper here
  • GEISSLER, PETER; TIMO HARTMANN & THY NEANG 2012. A new species of the genus Lygosoma Hardwicke & Gray, 1827 (Squamata: Scincidae) from northeastern Cambodia, with an updated identification key to the genus Lygosoma in mainland Southeast Asia. Zootaxa 3190: 56–68 - get paper here
  • Geissler, Peter; Truong Quang Nguyen, Trung My Phung, Robert Wayne Van Devender, Timo Hartmann, Balázs Farkas, Thomas Ziegler and Wolfgang Böhme 2011. A review of Indochinese skinks of the genus Lygosoma Hardwicke & Gray, 1827 (Squamata: Scincidae), with natural history notes and an identification key. BIOLOGIA 66 (6): 1159-1176 - get paper here
  • GRISMER, L. LEE; EVAN S. H. QUAH, ZAHARIL DUZULKAFLY & PAUL YAMBUN 2018. On the taxonomy of Lygosoma bampfyldei Bartlett, 1895 (Squamata: Scincidae) with descriptions of new species from Borneo and Peninsular Malaysia and the resurrection of Lygosoma schneideri Werner, 1900. Zootaxa 4438 (3): 528–550 - get paper here
  • Günther, A. 1864. The Reptiles of British India. London (Taylor & Francis), xxvii + 452 pp. - get paper here
  • Smith, M.A. 1914. Note on a rare lizard, (Lygosoma isodactylum). J. nat. Hist. Soc. Siam 1: 127-8
  • Smith, M.A. 1935. The fauna of British India, including Ceylon and Burma. Reptiles and Amphibia, Vol. II. Sauria. Taylor and Francis, London, 440 pp.
  • Taylor, E.H. 1963. The lizards of Thailand. Univ. Kansas Sci. Bull. 44: 687-1077. - get paper here
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