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Lygosoma malayana GRISMER, DZUKAFLY, MUIN, QUAH, KARIN, ANUAR & FREITAS, 2019

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Higher TaxaScincidae, Lygosominae, Scincoidea, Sauria, Squamata (lizards)
Subspecies 
Common NamesE: Malaysian Supple Skink 
SynonymSubdoluseps malayana GRISMER, DZUKAFLY, MUIN, QUAH, KARIN, ANUAR & FREITAS 2019
Lygosoma herberti — SWORDER 1933:103 (part)
Lygosoma herberti — GRISMER 2011:619
Lygosoma herberti — KARIN et al. 2018:361
Subdoluseps herberti — FREITAS et al. 2019: 22 (part) 
DistributionPeninsular Malaysia (Perak)

Type locality: Teluk Rubiah, Perak, Peninsular Malaysia (36.86694°N 10.466944°E WGS 84; 19 m in elevation).  
Reproduction 
TypesHolotype. LSUHC 10995, adult male collected by Zaharil Dzulkafly on 13 September 2012; Paratypes. Adult female ZRC 2.158 collected by G. Hope Sworder on 21 March 1929 from “Kampung Me- nora, Perak near Kuala Kangsar”, (as the specimen tag reads), Peninsular Malaysia (4.715275°N 100.944263°E; 40 m in elevation). Juvenile LSUHC 12098 collected by Mohd. A. Muin on 21 January 2013 from Bukit Panchor, Penang, Peninsular Malaysia (5.153889°N 100.54639°E; 79 m in elevation). 
DiagnosisDiagnosis. Grismer et al. 2019 consider Subdoluseps malayana sp. nov. to be a member of the genus Subduloseps in that it bears the suite of morphological characters that define the genus (Freitas et al. 2019). Subdoluseps malayana sp. nov. can be differentiated from all other Subdoluseps by having the combination of an adult SVL of 62.2–65.4 mm; six supralabials with the fourth being elongate; six infralabials; four supraoculars; frontoparietal contacting three supraoculars on each side; no postinterparietal; eight or nine superciliaries; 7–9 suboculars not separated by supralabials; prefrontals not in contact; one primary, two secondary, and three tertiary temporals, respectively; five or six nuchal scales; no deep postnasal groove; scaly lower eyelid lacking a window; two loreals; 29 or 30 transverse midbody scale rows bearing 3–5 keels; 57–64 paravertebral scale rows; 58–62 ventral scale rows; 19–21 caudal scale rows at the tenth subcaudal scale; seven or eight precloacal scales; smooth to weakly keeled subdigital finger lamellae; keeled to strongly keeled subdigital toe lamellae; 13–15 subdigital lamellae on fourth toe; and a bright yellow gular region, throat, and venter in adults. These characters are scored across S. herberti and S. samajaya in Table 4 (in Grismer et al. 2019).

Comparisons. Subdoluseps malayana sp. nov. is most closely related to S. samajaya but differs from it by having a statistically significant higher mean number of precloacal scales (7.3 versus 4.0), three versus four tertiary temporal scales, and the fourth as opposed to the fifth supralabial scale being elongate. Subdoluseps malayana sp. nov. differs from S. herberti by having a statistically significant higher mean number of paravertebral scales (61.0 versus 56.6) and ventral scales (61.0 versus 56.3) and by having a yellow versus a beige venter. Subdoluseps malayana sp. nov. differs from other Southeast Asia species such as Lygosoma angeli (Smith); L. anguinum (Theobald); L. bampfyldei Bartlett; S. bowringii (Günther) (usually); L. corpulentum Smith; L. boehmei Ziegler, Schmitz, Heidrich, Vu & Nguyen; S. frontoparietale (Taylor); L. haroldyoungi (Taylor); L. isodactylum (Günther); L. kinabatanganensis Grismer, Quah, Dzulkafly & Yambun; L. koratense Smith; L. peninsulare Grismer, Quah, Dzulkafly & Yambun; L. quadrupes (Linnaeus); L. schneideri Werner; L. tabonorum Heitz, Diesmos, Freitas, Ellsworth & Grismer; and L. veunsaiensis Geissler, Hartmann & Neang (usually); Riopa lineolata (Stolizcka); R. popae (Shreve); S. siamanensis (Siler, Heitz, Davis, Freitas, Aowphol, Termprayoon & Grismer) by having keeled as opposed to smooth scales. Subdoluseps malayana sp. nov. differs from L. opisthorhodum Werner by being larger (62.2–65.4 mm SVL versus 45 mm) and having a faint, light-colored dorsolateral stripe as opposed to a wide, well-defined, black dorsolateral stripe.
 
Comment 
EtymologyThe name malayana is in reference to this species being endemic to Peninsular Malaysia. 
References
  • GRISMER, L. LEE; ZAHARIL DZUKAFLY, MOHD ABDUL MUIN, EVAN S. H. QUAH, BENJAMIN R. KARIN, SHAHRUL ANUAR, ELYSE S. FREITAS 2019. A new skink of the genus Subdoluseps (Hardwicke & Gray, 1828) from Peninsular Malaysia. Zootaxa 4609 (2): 358–372 - get paper here
 
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