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Mantheyus phuwuanensis (MANTHEY & NABHITABHATA, 1991)

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Higher TaxaAgamidae (Draconinae), Sauria, Iguania, Squamata (lizards)
Common NamesE: Phuwua Rock Agama
G: Phuwua-Felsenagame 
SynonymPtyctolaemus phuwuanensis MANTHEY & NABHITABHATA 1991
Ptyctolaemus phuwuanensis — MANTHEY & SCHUSTER 1999: 101
Mantheyus phuwuanensis — ANANJEVA & STUART 2001
Mantheyus phuwuanensis — BARTS & WILMS 2003
Ptyctolaemus phuwuanensis — HALLERMANN & BÖHME 2003
Mantheyus phuwuanensis — SCHULTE et al 2004
Mantheyus phuwuanensis — MANTHEY 2010 
DistributionNE Thailand, WC Laos

Type locality: Phu Wua Wildlife Sanctuary, Nong Khai Province, Nong Dem Subdistrict, Bung Khan District, Thailand, 18°15' N, 103°58' E, 200-380 m elevation.  
TypesHolotype: ZMB 49040; paratype: ZMB 49041; additional specimens in ZMB, FMNH 
DiagnosisDiagnosis (genus). Femoral pores distinguish Mantheyus from all other genera of draconine agamids. The combination of femoral pores and haired skin sense organs distinguishes Mantheyus from all other agamids.

Comparisons: Ananjeva & Stuart 2001 compare Mantheyus phuwuanensis and Ptyctolaemus gularis in Table 1. 
CommentType species: Ptyctolaemus phuwuanensis MANTHEY & NABHITABHATA 1991
is the type species of the genus Mantheyus ANANJEVA & STUART 2001.

Ptyctolaemus phuwuanensis was assigned to the genus Ptyctolaemus Peters, 1864 because it shared the character of three parallel folds on each side of the throat with P. gularis. “Mantheyus phuwuanensis differs from Ptyctolaemus and all other draconines by the presence of femoral pores, and from all other agamids by having the combination of femoral pores and haired skin sense organs” (Ananjeva and Stuart 2001). 
EtymologyThe genus Mantheyus is named after Ulrich Manthey for his contributions to Southeast Asian herpetology. The specific epithet is derived from the name of the type locality. 
  • ANANJEVA, N. B. & STUART. B.L. 2001. The agamid lizard Ptyctolaemus phuwuanensis MANTHEY AND NABHITABHATA, 1991 from Thailand and Laos represents a new genus. Russ. J. Herpetol. 8 (3): 165-170 - get paper here
  • ANANJEVA, Natalia B.; Xianguang GUO and Yuezhao WANG 2011. Taxonomic Diversity of Agamid Lizards (Reptilia, Sauria, Acrodonta, Agamidae) from China: A Comparative Analysis. Asian Herpetological Research 2 (3): 117-128 - get paper here
  • Barts, M. & Wilms, T. 2003. Die Agamen der Welt. Draco 4 (14): 4-23 - get paper here
  • Chan-ard, T., Parr, J.W.K. & Nabhitabhata, J. 2015. A field guide to the reptiles of Thailand. Oxford University Press, NY, 352 pp. [see book reviews by Pauwels & Grismer 2015 and Hikida 2015 for corrections] - get paper here
  • Hallermann, J. & Böhme, W. 2003. A reinterpretation of the agamid genus Mantheyus Ananjeva and Stuart, 2001, as a junior synonym of Ptyctolaemus Peters, 1864, with comments on the problem of monotypic genera. Hamadryad 27 (2): 262-265
  • Manthey U 2010. Agamid Lizards of Southern Asia. Draconinae 2 -Leiolepidinae. Edition Chimaira, Terralog 7b, Frankfurt, 168 pp.
  • Manthey U; Nabhitabhata J 1991. Eine neue Agame, Ptyctolaemus phuwuanensis sp.n. (Sauria: Agamidae), aus Nordost-Thailand. Sauria 13 (4): 3-6 - get paper here
  • Manthey, S. & Manthey, U. 2005. Mantheyus phuwuanensis - from capture to captive breeding. Reptilia (GB) (42): 24-28 - get paper here
  • Manthey, S. & Manthey, U. 2005. Mantheyus phuwuanensis (MANTHEY & NABHITABHATA 1991) - vom Fang bis zur Nachzucht im Terrarium. Reptilia (Münster) 10 (51): 26-31 - get paper here
  • MANTHEY, U. & S. MANTHEY 2017. Amphibien und Reptilien von Laos – Ein Reisebericht – Teil 1: Phou Khao Khouay NBCA (Februar 1998). Sauria 39 (2): 35-55 - get paper here
  • Manthey,U. & SCHUSTER,N. 1999. Agamen, 2. Aufl. Natur und Tier Verlag (Münster), 120 pp. - get paper here
  • Schulte II, J. A., J. V. Vindum, H. Win, T. Thin, K. S. Lwin & A.K. Shein 2004. Phylogenetic relationships of the genus Ptyctolaemus (Squamata: Agamidae), with a description of a new species from Chin Hills of Western Myanmar. Proc. Cal. Acad. Sci., 55(12): 222-247 - get paper here
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