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Mediodactylus heteropholis (MINTON, ANDERSON & ANDERSON, 1970)

IUCN Red List - Mediodactylus heteropholis - Data Deficient, DD
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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common NamesE: Iraqi (Keel-scaled) Gecko 
SynonymTropiocolotes heteropholis MINTON, ANDERSON & ANDERSON 1970
Alsophylax persicus — REED & MARX 1959
Tropiocolotes heteropholis — MINTON et al. 1970: 357
Carinatogecko heteropholis — GOLUBEV & SZCZERBAK 1981
Carinatogecko heteropholis — ANDERSON 1999: 145
Carinatogecko heteropholis — RÖSLER 2000: 61
Mediodactylus heteropholis — ČERVENKA et al. 2010
Carinatogecko heteropholis — FATHINIA et al. 2011
Mediodactylus heteropholis — BAUER et al. 2013
Carinatogecko heteropholis — MOHAMMED et al. 2017 
DistributionNE Iraq (Erbil-Province), W Iran

Type locality: Pirman [= Primam] Hotel, Salahedin, Salahedin Nahiya, Iraq)  
Reproductionoviparous 
TypesHolotype: FMNH 74549 
DiagnosisDiagnosis: Mid-dorsal scales slightly larger than or subequal to abdominal scales; caudal tubercles distinctly swollen, but blunt, their posterior borders not upraised; scales of upper surface of foreann homogeneous; lower row of lateral subdigital scales acuminate; 15 subdigital lamellae on 4th toe (Anderson 1999: 145).


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CommentDistribution: See map in SMID et al. 2014 for distribution in Iran. 
EtymologyNamed after Greek heteros = different + Greek pholis = horny scale (Borror, 1988). 
References
  • Anderson, Steven C 1999. The lizards of Iran. Contributions to Herpetology Volume 15, Society for the Study of Amphibians and Reptiles, Saint Louis, Missouri: i-vii, 1-442 [review in Copeia 2000 (4): 1144] - get paper here
  • BAUER, AARON M.; RAFAQAT MASROOR, JAMES TITUS-MCQUILLAN, MATTHEW P. HEINICKE,, JUAN D. DAZA & TODD R. JACKMAN 2013. A preliminary phylogeny of the Palearctic naked-toed geckos (Reptilia: Squamata: Gekkonidae) with taxonomic implications. Zootaxa 3599 (4): 301–324 - get paper here
  • ČERVENKA, JAN; DANIEL FRYNTA & LUKÁŠ KRATOCHVÍL 2010. Phylogenetic relationships of the gecko genus Carinatogecko (Reptilia: Gekkonidae). Zootaxa 2636: 59–64 - get paper here
  • Fathinia, B., Rastegar-Pouyani, N., And Mohamadi, H. 2011. Sexual dimorphism in Carinatogecko heteropholis (Minton, Anderson, and Anderson, 1970) (Sauria: Gekkonidae) from Ilam Province, western Iran. Amphibian & Reptile Conservation 5 (1): 47-53 - get paper here
  • Fathinia, Behzad; Rasoul Karamiani, Hamid Darvishnia, Naghi Heidari, and Nasrullah Rastegar-Pouyani. 2011. A new species of Carinatogecko (Sauria: Gekkonidae) from Ilam Province, western Iran. Amphibian & Reptile Conservation 5 (1): 61-74 - get paper here
  • Golubev, M. L., and N. N. Szczerbak 1981. Carinatogecko gen. n. (Reptilia, Gekkonidae) - a new genus from south-west Asia [in Russian]. Vestnik Zoologii, Kiev 1981 (5):34—41.
  • Kamali, Kamran 2020. A Field Guide to the Reptiles and Amphibians of Iran. Edition Chimaira, Frankfurt am Main, Germany (www.chimaira.de). 574 pp.
  • Leviton,A.E.; Anderson,S.C.; Adler, K.; Minton,S.A. 1992. Handbook to Middle East Amphibians and Reptiles. SSAR, Oxford, Ohio (Contr. to Herpetol. No. 8), 1-252
  • Minton, S. A., Jr., S. C. Anderson, and J. A. Anderson. 1970. Remarks on some geckos from southwest Asia, with descriptions of three new forms and a key to the genus Tropiocolotes. Proc. Cal. Acad. Sci. 37: 333—362 - get paper here
  • Mohammed, Rihab Ghaleb; Fadhel Aabbas Rhadi, Nasrullah Rastegar-Pouyani, Eskandar Rastegar-Pouyani, Seyyed Saeed Hosseinian Yousefkhani 2017. Zoogeography of Lizards Fauna from Central and Southern Iraq with a Checklist of Iraqi Lizard’s Fauna. Russ. J. Herpetol. 24 (3): 193-201 - get paper here
  • RASTEGAR-POUYANI, NASRULLAH; HAJI GHOLI KAMI, MEHDI RAJABZADEH, SOHEILA SHAFIEI AND STEVEN CLEMENT ANDERSON 2008. Annotated Checklist of Amphibians and Reptiles of Iran. Iranian Journal of Animal Biosystematics 4 (1): 7-30
  • Reed, Charles A. & Marx, Hymen 1959. A herpetological collection from Northeastern Iraq. Transactions of the Kansas Academy of Science 62 (1): 91-122 - get paper here
  • Rösler, H. 2000. Kommentierte Liste der rezent, subrezent und fossil bekannten Geckotaxa (Reptilia: Gekkonomorpha). Gekkota 2: 28-153
  • Shcherbak, N. N. [= Szczerbak], and M. L. Golubev 1996. Gecko fauna of the USSR and contiguous regions. SSAR [Russian original published in 1986]. [book review in Salamandra 36 (2): 138]
  • Sindaco, R. & Jeremcenko, V.K. 2008. The reptiles of the Western Palearctic. Edizioni Belvedere, Latina (Italy), 579 pp. - get paper here
  • ŠMÍD, JIŘÍ; JIŘÍ MORAVEC, PETR KODYM, LUKÁŠ KRATOCHVÍL, SEYYED SAEED HOSSEINIAN YOUSEFKHANI, ESKANDAR RASTEGAR-POUYANI & DANIEL FRYNTA 2014. Annotated checklist and distribution of the lizards of Iran. Zootaxa 3855 (1): 001–097 - get paper here
  • Torki, F. 2011. Description of a new species of Carinatogecko (Squamata: Gekkonidae) from Iran. Salamandra 47 (2): 103-111 - get paper here
  • Torki, Farhang 2012. Notes on Sexual Dimorphism of Carinatogecko heteropholis (Reptilia: Gekkonidae) (Minton, Anderson et Anderson, 1970). Russ. J. Herpetol. 19 (4): 284-286 - get paper here
 
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