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Mediodactylus heterocercus (BLANFORD, 1874)

IUCN Red List - Mediodactylus heterocercus - Least Concern, LC
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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
SubspeciesMediodactylus heterocercus heterocercus (BLANFORD 1874)
Mediodactylus heterocercus mardinensis (MERTENS 1924) 
Common NamesE: Asia Minor Thin-Toed Gecko 
SynonymGymnodactylus heterocercus BLANFORD 1874: 453
Gymnodactylus heterocercus — BOULENGER 1885: 30
Gymnodactylus heterocercus — WERMUTH 1965: 54
Cyrtopodion (Mediodactylus) heterocercus — BÖHME 1985: 96
Mediodactylus heterocercus — KLUGE 1993
Cyrtopodion heterocercum — ANDERSON 1999
Cyrtopodion (Mediodactylus) heterocercus — RÖSLER 2000: 69
Cyrtopodion heterocercum — FROST 2007 (pers. comm.)
Cyrtopodion (Mediodactylus) heterocercum — SINDACO & JEREMČENKO 2008
Mediodactylus heterocercus — BAUER et al. 2013
Mediodactylus heterocercum — ŠMÍD et al. 2014
Mediodactylus heterocercum — MOZAFFARI et al. 2016

Mediodactylus heterocercus mardinensis (MERTENS 1924)
Gymnodactylus heterocercus mardinensis MERTENS 1924
Cytodactylus hetrocercus [sic] — NADER & JAWDAT 1976
Cyrtopodion heterocercus — KLUGE 1985
Tenuidactylus heterocercus mardinensis — SZCZERBAK & GOLUBEV 1986
Cyrtopodion (Mediodactylus) heterocercus mardinensis — LEVITON et al. 1992
Cyrtopodion mardinensis — RÖSLER 2000
Cyrtopodion heterocercus mardinensis — UGURTAS et al. 2007
Cyrtopodion heterocercum mardinense — SINDACO & JEREMČENKO 2008
Mediodactylus heterocercus mardinensis — BAUER et al. 2013
Cyrtpodion [sic] heterocercus mardinensis — MOHAMMED et al. 2017 
DistributionIran (Kermanshahan), SE Turkey (Mardin, Anatolia), S Iraq, Syria

heterocercus: Iran etc.; Type locality: Hamandran, West Persia [= Hamadan, Iran]

mardinense: Turkey (SE Anatolia, N Syria, N Iraq  
Reproductionoviparous 
TypesHolotype: MRSN (= MSNTO)
Lectotype: TZM R2532, adult female, “Persia dono Doria”, designated by SZCZERBAK & GOLUBEV (1986:183)]
Holotype: SMF 8198 [mardinensis] 
DiagnosisDiagnosis: Dorsum with enlarged trihedral tubercles in 12-14 longitudinal rows; no enlarged subcaudal plates, tail covered below with strongly keeled imbricate scales in 5-7 longitudinal rows; sides oftail with greatly enlarged sharply keeled mucronate scales (Fig. 72C, Anderson 1999: 158). 
CommentNot listed by Engelmann et al. (1993).

Distribution: See map in SMID et al. 2014 for distribution in Iran. 
References
  • Afrasiab, Saman R. and Sarbaz I. Mohamad 2009. A study on cave-dwelling geckos in Iraq, with the description of a new species from Saffine mountain (Reptilia: Gekkonidae). Zoology in the Middle East 47:49-56 - get paper here
  • Akman, B.; M. Z. Yildiz et al. 2018. On the herpetofauna of the East Anatolian Province of Bitlis (Turkey) (Amphibia; Reptilia). Herpetozoa 31 (1/2): 69 - 82 - get paper here
  • Anderson, Steven C 1999. The lizards of Iran. Contributions to Herpetology Volume 15, Society for the Study of Amphibians and Reptiles, Saint Louis, Missouri: i-vii, 1-442 [review in Copeia 2000 (4): 1144] - get paper here
  • BAUER, AARON M.; RAFAQAT MASROOR, JAMES TITUS-MCQUILLAN, MATTHEW P. HEINICKE,, JUAN D. DAZA & TODD R. JACKMAN 2013. A preliminary phylogeny of the Palearctic naked-toed geckos (Reptilia: Squamata: Gekkonidae) with taxonomic implications. Zootaxa 3599 (4): 301–324 - get paper here
  • Bird, C. G. 1936. The distribution of reptiles and amphibians in Asiatic Turkey with notes on a collection from the Vilayets of Adana, Gaziantep and Malatya. Ann. Mag. Nat. Hist. (10) 18: 257-281. - get paper here
  • Blanford, W.T. 1874. Descriptions of new Reptilia and Amphibia from Persia and Baluchistan. Ann. Mag. nat. Hist. (4) 14: 31-35 - get paper here
  • Böhme, W. 1985. Zur Nomenklatur der paläarktischen Bogenfingergeckos, Gattung Tenuidactylus Szczerbak & Golubev 1984 (Reptilia: Gekkonidae). Bonner Zoologische Beiträge 36: 95-98 - get paper here
  • Boulenger, G.A. 1885. Catalogue of the lizards in the British Museum (Nat. Hist.) I. Geckonidae, Eublepharidae, Uroplatidae, Pygopodidae, Agamidae. London: 450 pp. - get paper here
  • Kamali, Kamran 2020. A Field Guide to the Reptiles and Amphibians of Iran. Edition Chimaira, Frankfurt am Main, Germany (www.chimaira.de). 574 pp.
  • Leviton,A.E.; Anderson,S.C.; Adler, K.; Minton,S.A. 1992. Handbook to Middle East Amphibians and Reptiles. SSAR, Oxford, Ohio (Contr. to Herpetol. No. 8), 1-252
  • Mertens, R. 1967. Die herpetologische Sektion des Natur-Museums und Forschungs-Institutes Senckenberg in Frankfurt am Main nebst einem Verzeichnis ihrer Typen. Senckenbergiana Biologica 48: 1-106 - get paper here
  • Mertens, Robert 1924. Ein neuer Gecko aus Mesopotamien. Senckenbergiana 6 (1-2): 84 - get paper here
  • Mohammed, Rihab Ghaleb; Fadhel Aabbas Rhadi, Nasrullah Rastegar-Pouyani, Eskandar Rastegar-Pouyani, Seyyed Saeed Hosseinian Yousefkhani 2017. Zoogeography of Lizards Fauna from Central and Southern Iraq with a Checklist of Iraqi Lizard’s Fauna. Russ. J. Herpetol. 24 (3): 193-201 - get paper here
  • Moravec, J.; Modry, D. 1994. On the occurrence of Cyrtodactylus heterocercus mardinensis and Pseudocerastes persicus fieldi in Syria. Zoology in the Middle East 10: 53-56 - get paper here
  • Mozaffari, Omid; Kamran Kamali and Hadi Fahimi 2016. The Atlas of Reptiles of Iran. Iran Department of the Environment, 362 pp. [2014]
  • RASTEGAR-POUYANI, NASRULLAH; HAJI GHOLI KAMI, MEHDI RAJABZADEH, SOHEILA SHAFIEI AND STEVEN CLEMENT ANDERSON 2008. Annotated Checklist of Amphibians and Reptiles of Iran. Iranian Journal of Animal Biosystematics 4 (1): 7-30
  • Rastegar-Pouyani, Nasrullah; Hamzeh Oraei, and Morteza Johari 2009. New Records of the Geckonid Lizard, Cyrtopodion heterocercum heterocercum (Sauria: Gekkonidae) from Isfahan Province, Central Iran, with an Extended Description and Notes on Distribution. Russ. J. Herpetol. 16 (2): 95-100 - get paper here
  • Rösler, H. 2000. Kommentierte Liste der rezent, subrezent und fossil bekannten Geckotaxa (Reptilia: Gekkonomorpha). Gekkota 2: 28-153
  • Rösler, H. 2000. Die postanale Beschuppung bei Cyrtodactylus GRAY 1827 und Cyrtopodion FITZINGER 1843 - funktionelle und taxonomische Aspekte (Sauria: Gekkonidae). Gekkota 2: 154-207
  • RÖSLER, H. &; SCHMIDTLER, J. F. &; MORAVEC, J. 2012. Bemerkungen zu einigen Unterarten von Mediodactylus kotschyi (STEINDACHNER, 1870) und Mediodactylus heterocercus mardinensis (MERTENS, 1924) in der südlichen Türkei und in angrenzenden Gebieten. Herpetozoa 25 (1/2): 25-46 - get paper here
  • SAFAEI-MAHROO, BARBOD; HANYEH GHAFFARI, & STEVEN C. ANDERSON 2016. A new genus and species of gekkonid lizard (Squamata: Gekkota: Gekkonidae) from Hormozgan Province with a revised key to gekkonid genera of Iran. Zootaxa 4109 (4): 428–444 - get paper here
  • Schleich, H. H. 1977. Distributional maps of reptiles of Iran. Herpetological Review 8: 126-129. - get paper here
  • Shcherbak, N. N. [= Szczerbak], and M. L. Golubev 1996. Gecko fauna of the USSR and contiguous regions. SSAR [Russian original published in 1986]. [book review in Salamandra 36 (2): 138]
  • Sindaco, R. & Jeremcenko, V.K. 2008. The reptiles of the Western Palearctic. Edizioni Belvedere, Latina (Italy), 579 pp. - get paper here
  • Sindaco, Roberto; Alberto Venchi, Giuseppe M. Carpaneto, Marco A. Bologna 2000. The Reptiles of Anatolia: a Checklist and Zoogeographical analysis. Biogeographia, 21-2000: 441-554. - get paper here
  • ŠMÍD, JIŘÍ; JIŘÍ MORAVEC, PETR KODYM, LUKÁŠ KRATOCHVÍL, SEYYED SAEED HOSSEINIAN YOUSEFKHANI, ESKANDAR RASTEGAR-POUYANI & DANIEL FRYNTA 2014. Annotated checklist and distribution of the lizards of Iran. Zootaxa 3855 (1): 001–097 - get paper here
  • Torki, F. 2011. Description of a new species of Carinatogecko (Squamata: Gekkonidae) from Iran. Salamandra 47 (2): 103-111 - get paper here
  • Uca, Onur; Hüseyin Arikan, Kerim Cicek 2017. Blood cell morphology of Turkish gekkonid lizards (Squamata: Sauria: Gekkonidae, Phyllodactylidae). Herpetozoa 30 (1/2): 29 - 37 - get paper here
  • UGURTAS¸ Ismail HAKKI; HIKMET SAMI YILDIRIMHAN & Murat Sevinc 2007. Distribution of the Gekkonidae Species in Southeast Anatolia, Turkey, and New Localities. Turkish Journal of Zoology 31: 137-141 - get paper here
 
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