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Meroles squamulosus (PETERS, 1854)

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Higher TaxaLacertidae, Eremiadinae, Sauria, Lacertoidea, Squamata (lizards)
Subspecies 
Common NamesE: Savanna Lizard, Common Rough-scaled (Sand) Lizard 
SynonymIchnotropis squamulosa PETERS 1854: 617
Ichnotropis squamulosa — FITZSIMONS 1943: 350
Ichnotropis squamulosa — HUEY & PIANKA 1981
Ichnotropis squamulosa — AUERBACH 1987: 130
Ichnotropis squamulosa — BROADLEY & HOWELL 1991: 18
Ichnotropis squamulosa — PIETERSEN et al. 2013
Meroles squamulosa — ENGLEDER et al. 2013
Meroles squamulosa — EDWARDS et al. 2013
Meroles squamulosus — SPAWLS et al 2018: 20
Meroles squamulosus — SPAWLS et al 2018: 203 
DistributionTanzania, Zambia, S Angola, Zimbabwe, Botswana, Namibia, N South Africa, Malawi.

Type locality: Tete, Mozambique  
Reproductionoviparous 
TypesSyntypes: ZMB 6614, 6615, 6616 
Diagnosis 
CommentSynonymy: EDWARDS et al. 2013 proposed the name Meroles squamulosa as “comb. nov.” but Engleder’s paper was published earlier. 
References
  • Auerbach, R.D. 1987. The Amphibians and Reptiles of Botswana. Mokwepa Consultants, Botswana, 295 pp.
  • Baeckens, Simon, Jan Scholliers and Raoul Van Damme. 2016. Geographic Distribution: Meroles squamulosus (Common Rough-scaled lizard). Herpetological Review 47 (1): 81 - get paper here
  • Baptista NL, António T, Branch WR. 2019. The herpetofauna of Bicuar National Park and surroundings, southwestern Angola: a preliminary checklist. Amphibian & Reptile Conservation 13(2) [Special Section]: 96–130 (e203) - get paper here
  • Bates, M.F.; Branch, W.R., Bauer, A.M.; Burger, M., Marais, J.; Alexander, G.J. & de Villliers, M.S. (eds.) 2014. Atlas and Red List of the Reptiles of South Africa, Lesotho, and Swaziland. Suricata 1. South African National Biodiversity Institute, Pretoria, 512 pp.
  • Bauer, Aaron M.;Günther, Rainer 1995. An annotated type catalogue of the lacertids (Reptilia: Lacertidae) in the Zoological Museum, Berlin. Mitt. Zool. Mus. Berlin 71 (1): 37-62 - get paper here
  • Bittencourt-Silva GB. 2019. Herpetological survey of western Zambia. Amphibian & Reptile Conservation 13(2) [Special Section]: 1-28 (e181) - get paper here
  • Branch, William R. 1993. A Photographic Guide to Snakes and Other Reptiles of Southern Africa. Cape Town: Struik Publishers, 144 S.
  • Broadley, D. G. & HOWELL, K. M. 1991. A check list of the reptiles of Tanzania, with synoptic keys. Syntarsus 1: 1—70
  • Broadley, D.G. 1962. On some reptile collections from the North-Western and North-Eastern Districts of Southern Rhodesia 1958-1961, with descriptions of four new lizards. Occ. Pap. Nat. Mus. South. Rhodesia 26 (B): 787-843
  • Buruwate, T. C., & Lloyd-Jones, D. J. 2024. Amphibian and Reptile Diversity of Niassa Special Reserve, Northern Mozambique. Journal of East African Natural History, 113(1), 1-18 - get paper here
  • Conradie W and Branch WR. 2016. The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian & Reptile Conservation 10 (2) [Special Section]: 6–36 - get paper here
  • Conradie W, Keates C, Verburgt L, Baptista NL, Harvey J, Júlio T, Neef G. 2022. Contributions to the herpetofauna of the Angolan Okavango-Cuando-Zambezi River drainages. Part 2: Lizards (Sauria), chelonians, and crocodiles. Amphibian & Reptile Conservation 16(2): 181–214 (e322) - get paper here
  • CONRADIE, W., DOUCETTE-RIISE, S., VANHOOYDONCK, B., ENGELBRECHT, H., MEASEY, G.J., & TOLLEY, K. 2011. Herpetological survey of Rooipoort Nature Reserve, Northern Cape, South Africa. African Herp News (53): 35-41 - get paper here
  • CONRADIE, W., ENGELBRECHT, H., HERREL, A., MEASEY, G.J., NIELSEN, S.V., V ANHOOYDONCK, B., & TOLLEY, K.A. 2011. Reptile survey of Venetia Limpopo Nature Reserve, Limpopo Province - South Africa. African Herp News (55): 6-12 - get paper here
  • EDWARDS, SHELLEY; WILLIAM R. BRANCH, BIEKE VANHOOYDONCK, ANTHONY HERREL, G. JOHN MEASEY, KRYSTAL A. TOLLEY 2013. Taxonomic adjustments in the systematics of the southern African lacertid lizards (Sauria: Lacertidae). Zootaxa 3669 (2): 101–114 - get paper here
  • Engleder, A., Haring, E., Kirchhof, S. and Mayer, W. 2013. Multiple nuclear and mitochondrial DNA sequences provide new insights into the phylogeny of South African Lacertids (Lacertidae, Eremiadinae). Journal of Zoological Systematics and Evolutionary Research, 51: 132–143. doi: 10.1111/jzs.12012 - get paper here
  • FitzSimons, V.F. 1943. The lizards of South Africa. Transvaal Museum Memoir No.1 (Pretoria), 528 pp.
  • Herrmann, H.-W.; W.R. Branch 2013. Fifty years of herpetological research in the Namib Desert and Namibia with an updated and annotated species checklist. Journal of Arid Environments 93: 94–115 - get paper here
  • Huey, Raymound B.; Eric R. Pianka 1981. Ecological Consequences of Foraging Mode. Ecology 62 (4):991-999
  • Jacobsen, Niels H.G.; Errol W. Pietersen & Darren W. Pietersen 2010. A preliminary herpetological survey of the Vilanculos Coastal Wildlife Sanctuary on the San Sebastian Peninsula, Vilankulo, Mozambique. Herpetology Notes 3: 181-193
  • Kirchhof, S.; Engleder, A.; Mayer, W. & Richter, K. 2011. Die Radiation der Lacertiden des südlichen Afrikas. Elaphe 19 (4): 6-11
  • Loveridge, A. 1957. On a third collection of reptiles taken in Tanganyika by C.J.P. Ionides, Esq. Tanganyika Notes and Records 43: 1-19
  • Marques, Mariana P.; Luis M. P. Ceríaco , David C. Blackburn , and Aaron M. Bauer 2018. Diversity and Distribution of the Amphibians and Terrestrial Reptiles of Angola -- Atlas of Historical and Bibliographic Records (1840–2017). Proc. Cal. Acad. Sci. (Ser. 4) 65: 1-501 (Supplement II)
  • Miguel, A. R., Kaiser, H., Vargas, N. D., & Borges-Martins, M. 2024. Diversity and natural history of reptiles from coastal environments in Gilé National Park, northern Mozambique. Herpetology Notes, 17, 173-199
  • Peters,W.C.H. 1854. Diagnosen neuer Batrachier, welche zusammen mit der früher (24. Juli und 17. August) gegebenen Übersicht der Schlangen und Eidechsen mitgetheilt werden. Ber. Bekanntmach. Geeignet. Verhandl. Königl.-Preuss. Akad. Wiss. Berlin 1854: 614-628
  • Pianka, Eric R. 1971. Lizard species density in the Kalahari Desert. Ecology 52 (6): 1024-1029 - get paper here
  • Pietersen DW, Pietersen EW, Conradie W. 2017. Preliminary herpetological survey of Ngonye Falls and surrounding regions in south-western Zambia. Amphibian & Reptile Conservation 11(1) [Special Section]: 24–43 (e148 - get paper here
  • Pietersen, Darren W. Pietersen, Errol W. Haacke, Wulf D. 2013. First herpetological appraisal of the Parque Nacional de Banhine, Gaza Province, southern Mozambique. Annals of the Ditsong National Museum of Natural History 3: 153-163 - get paper here
  • Pietersen, Darren, Verburgt, Luke & Davies, John 2021. Snakes and other reptiles of Zambia and Malawi. Struik Nature / Penguin Random House South Africa, 376 pp., ISBN 9781775847373
  • Schmidt, A. 2001. Autökologie und Fortpflanzungsbiologie von Ichnotropis squamulosa PETERS 1854 (Reptilia: Lacertidae). Salamandra 37 (3): 157-174 - get paper here
  • Schmidt, A. D. 2005. Habitatnutzung, Raumbedarf und Mobilität von Ichnotropis squamulosa PETERS 1854 (Reptilia, Lacertidae) in der Trockensavanne Südafrikas. Sauria 27 (4): 3-14 - get paper here
  • SCHMIDT, A.D. 2006. Zur Populationsökologie von Ichnotropis squamulosa PETERS, 1854 (Reptilia, Lacertidae) in der Trockensavanne Südafrikas. Sauria 28 (1): 31-38 - get paper here
  • Spawls, S.; Howell, K.; Drewes, R.C. & Ashe, J. 2002. A field guide to the reptiles of East Africa. Academic Press, 543 pp. [reviews in HR 34: 396 and Afr. J. Herp. 51; 147] - get paper here
  • Spawls, Steve; Kim Howell, Harald Hinkel, Michele Menegon 2018. Field Guide to East African Reptiles. Bloomsbury, 624 pp. - get paper here
  • Tolley KA, Telford NS, Makhubo BG, Power RJ, Alexander GJ 2023. Filling the gap: Noteworthy herpetological discoveries in North West Province, South Africa. Zoosystematics and Evolution 99(1): 101-116 - get paper here
 
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