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Nactus aktites ZUG, 2020

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common NamesE: Madang Coastal Slender-toed Gecko 
SynonymNactus aktites ZUG 2020: 45 
DistributionN Papua New Guinea (from Wewak, East Sepik Province, to Alexishafen, Madang Province)

Type locality: Papua New Guinea, Madang Province, Awar [14.1362° 144.8664°]  
Reproduction 
TypesHolotype. IRSNB 15875.1516, adult male, collected by Benoit Mys in early 1980s.
Paratypes. IRSNB 15875.1500, 15875.1570, 15875.1618, 15875.1649, adult males, and IRSNB 15875.1526, 15875.1547, 15875.1569, 15875.1621, adult females with the same collection data as the holotype. 
DiagnosisDefinition. A bisexual taxon of geckos (Gekkoni- nae) with moderate adults (♀♀ 49–61 mm, ♂♂ 44–58 mm SVL) with keeled subcaudals, large postmental scales (median 15), no or slight postmental-infralabial (0–2) contact, moderate number of dorsal tubercle rows (14–20) and moderately high number of tubercles (26–33) in parasagittal row (TubRow), enlarged tubercule only on dorsal surface of crus, and no or few precloa- cal pores (0–7) in males and usually none in females. Diagnostic summary in Table 8.

Variation. Females and males display slight met- ric differences (adult ♀♀ 54.2, 46.8–60.8 mm; adult ♂♂ 51.3, 43.8–58.0 mm SVL) and share similar proportions (total adult sample): TrunkL/SVL 39.5%, 35%–45%; HeadL/SVL 27.6%, 23%–30; HeadW/SVL 19.6%, 16%–22%; HeadW/ HeadL 71.1%, 60%–80%; EyeD/SVL 6.0%, 5%–8%; EyeD/ HeadL 21.7%, 18%–27%; NarEye/HeadL 31.3%, 28%–37%; Interorb/HeadL 24.9%, 17%–33%; SnW/HeadL 13.5%, 12%–16%.
Typical head scale pattern of small, granular scales except for those bordering nares and mouth. Snout with large ros- tral scale bearing middorsal cleft posteriorly; large supralabi- als (3, 3–4) and infralabials (3, 2–4) in front of anterior edge of orbit, first supra- and infralabial largest of respective series. Chin with large trapezoidal mental and usually moderately large postmentals (Postm = 15, 0–22) and usually no genial scale between postmental and first surpalabial (0, 0–2 PmLab); dorsally, trunk with 18, 14–20 (DorsTub) enlarged tubercles transversely on dorsum and with 29, 26–33 tubercles (TubRow) longitudinally along trunk; 10, 8–11 tubercle rows trans- versely between hindlimbs (TubHip). Uniform covering of small tuberculate-like scale on dorsal half of forelimb and tubercles only on crus (TubHindl = 1). Tail with small uniform scales dor- sally and laterally, ventrally with uniform, small keeled scales (Subcaud = 0). Usually, single cloacal spurs (1, 1–2 CloacS) on each side. Ventral scales from chin to vent small, granular, some- what larger on chest and abdomen. Precloacal pores, males 0–7, median 4; females 0–7, median 0. Forefoot with narrow digits, their lengths usually 3≈4>2≈5>1; 17, 14–20 lamellae beneath fourth digit (4FingLm), 5, 4–6 scale rows on palm; hindfoot with narrow digits, their lengths 4≈3>2≈5>1; 21, 19–20 lamel- lae beneath fourth toe (4ToeLm).

Coloration in life not reported. In alcohol, not recorded. 
CommentDistribution: see map in Zug 2020: 47 (Fig. 18). 
EtymologyThe Greek aktites is a masculine noun for a shore or coast dweller. The name reflects this species’ pres- ence on coastal islands and coastal lowlands of Madang and East Sepik Provinces. The specific name is a noun in apposition. 
References
  • Zug, George R. 2020. Diversity in Pacific Slender-Toed Geckos, Nactus pelagicus Complex (Reptilia: Squamata), of New Guinea and Adjacent Islands. Smithsonian Contributions to Zoology, (651): x + 92 pages - get paper here
 
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