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Nactus arfakianus (MEYER, 1874)

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Common NamesE: Arafak Slender-toed Gecko 
SynonymGymnodactylus (Heteronota) arfakianus MEYER 1874: 129
Nactus arfakianus — ZUG 2020: 39 
DistributionIndonesia (Schouten Islands in the mouth of Cenderawasih Bay)

Type locality: “Neu-Guinea”; subsequently, Meyer (1887) identified the locality as Doré, a town on the northeastern coast of Pulau Yapen (=Kepulauan Yapen)  
ReproductionNactus pelagicus is a composite of unisexual [parthenogenetic] and bisexual populations. Note however that Zug and Moon (1995) recognized Nactus pelagicus as the asexual species for the populations in southern Vanuatu and elsewhere in Oceania, and N. multicarinatus for the bisexual species from Vanuatu and the southern Solomon Islands. 
TypesHolotype: lost (destroyed during the Dresden firebombing in February 1945. Zug 2020 has not designated a neotype because he has not located any specimens from Doré or Yapen.
Other specimens: BPBM 3951, Biak Island; UMMZ 122449, (Numfoor Island) are from the Schouten Islands, of which Yapen is a member. 
DiagnosisDefinition. A bisexual taxon of geckos (Gekkoninae) of moderate adults (47–52 mm SVL) with keeled subcaudals, moderate to large postmental scales (12–21), no postmentalinfralabial (PmLab) contact, moderate number of dorsal tubercle rows (15–18) and moderate number of tubercles (29–34) in parasagittal row (TubRow), tubercles on dorsal surface of thigh and crus, and 15 precloacal pores in a female (Figure 13). Diagnostic summary in Table 8.
Coloration in life is unknown. In alcohol, dorsal background medium-dark brown with series of paired dark-brown spots/blotches extending from anterior neck onto base of tail; head medium-dark brown with light-brown edging of supraand infralabials; venter from chin to base of tail uniform brown (this description is not based on any type material but from the other two specimens that were available to Zug 2020 and listed under types). 
CommentSynonymy: this species has been considered as a synonym of N. pelagicus for most of recent history until it was revalidated by Zug 2020 based on 2 specimens. 
Etymology“Arfak” derives from the language of the coastal Biak people of the Vogelkop Peninsula and appears to be used for their lowland area (“inferior”) relative to the adjacent Arfak Mountains, the highest mountains on the Vogelkop. Apparently, Meyer used Arfak in a broader geographic sense to label the entire region. 
  • Meyer, A.B. 1874. Eine Mittheilung von Hern. Dr. Adolf Berhard Meyer über die von ihm auf Neu-Guinea und den Inseln Jobi, Mysore und Mafoor im Jahre 1873 gesammelten Amphibien. Monatsber. K. Preuss. Akad. Wiss. Berlin 1874: 128-140 - get paper here
  • Zug, George R. 2020. Diversity in Pacific Slender-Toed Geckos, Nactus pelagicus Complex (Reptilia: Squamata), of New Guinea and Adjacent Islands. Smithsonian Contributions to Zoology, (651): x + 92 pages - get paper here
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