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Nactus erugatus ZUG, 2020

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Common NamesE: Milne Bay Smooth-tailed Slender-toed Gecko 
SynonymNactus erugatus ZUG 2020: 62 
DistributionPapua New Guinea (Milne Bay Province: SE Owen Stanley Range)

Type locality: Duabo, Pini Range, 10°06′S 151°15′E, 300 m, Milne Bay Province, Papua New Guinea.  
TypesHolotype. BPBM 15448, adult male, collected by Fred Kraus on 1 May 2002 (Figure 29).
Paratypes. Adult females, BPBM 15449, with same collecting data as holotype, and BPBM 15451–452 from Owen Stanley Range, Mount Pekopekowana, Garden Camp, 10°17.104′S, 150°10.930′E, 330 m, Milne Bay Province, Papua New Guinea, collected by Fred Kraus and I. Bigilale on 7 May 2002; adult males, BPBM 15443 from Bwaona River, Alotau, 10°30′14′′S, 150°18′50′′E, 90 m, Milne Bay Province, Papua New Guinea, collected by Fred Kraus on 7 April 2002, and BPBM 15450 from Mount Pekopekowana, Garden Camp, 10°17.104′S, 150°10.930′E, 330 m, Milne Bay Province, Papua New Guinea, collected by Fred Kraus and I. Bigilale on 7 May 2002. 
DiagnosisDefinition. A bisexual taxon of geckos (Gekkoni- nae) with large adults (♀♀ 66–72 mm, ♂♂ 59–69 mm SVL) with smooth subcaudals, moderate-sized postmental scales (10–20, median 14), no or slight postmental-infralabial (PmLab = 0–1, median 0) contact, moderate number of dorsal tubercle rows (15–18) and moderate number of tubercles (28–34, usually ≥29) in parasagittal row (TubRow), tubercles usually present on dorsal surface of thigh and crus, and precloacal pores 12–14 in males and 6–12 in females. Diagnostic summary in Table 8.

Variation. Females and males display slight metric differences (adult ♀♀ 69.7, 66.2–72.2 mm; adult ♂♂ 63.1, 59.0– 69.2 mm SVL), although sharing similar proportions (total adult sample): TrunkL/SVL 41.6%, 38%–45%; HeadL/SVL 27.5%, 26%–28%; HeadW/SVL 17.6%, 16%–18%; HeadW/HeadL 63.9%, 62%–66%; EyeD/SVL 6.9%, 6%–7%; EyeD/HeadL 25.0%, 23%–25%; NarEye/HeadL 32.3%, 31%–33%; Inter- orb/HeadL 25.6%, 23%–27%; SnW/HeadL 14.5%, 14%–15%.
Typical head scale pattern of small, granular scales except for those bordering nares and mouth. Snout with large rostral scale bearing middorsal cleft posteriorly; supranasals moderate sized, usually in contact on midline or separated by small granu- lar scale (internasal); large supralabials (4, 3–4) and infralabi- als (3, 3–4) in front of anterior edge of orbit, first supra- and infralabial largest of respective series. Chin with large trapezoi- dal mental and small to moderate postmentals (Postm = 14, 10–20) and usually no genial scale between postmental and first supralabial (0, 0–1 PmLab); dorsally, trunk with 17, 15–18 (DorsTub) enlarged tubercles transversely on dorsum and with 30, 28–34 tubercles (TubRow) longitudinally along trunk; 10, 8–10 tubercle rows transversely between hindlimbs (TubHip). Enlarged tubercles on crus and femur (TubHindl = 2). Tail with small uniform scales dorsally and laterally, ventrally typically with uniform, small smooth scales (Subcaud = 0). Usually, pair of cloacal spurs (2, 1–2 CloacS) on each side. Ventral scales from chin to vent small, granular, somewhat larger on chest and abdo- men. Precloacal pores, males 12–14, females 6–12. Forefoot with narrow digits, their lengths usually 3≈4>2≈5>1; 18, 17–19 lamel- lae beneath fourth digit (4FingLm), 5, 4–7 scale rows on palm; hindfoot with narrow digits, their lengths 3≈4>2≈5>1; 22, 19–23 lamellae beneath fourth toe (4ToeLm).
Coloration in life not reported. In alcohol, color and pattern match the general description of the holotype. 
CommentDistribution: see map in Zug 2020: 53 (Fig. 22). 
EtymologyThe specific name erugatus derives from the Latin erugo, “to take wrinkles from or smooth,” and refers to the smooth or keel-free subcaudals of this taxon. 
  • Zug, George R. 2020. Diversity in Pacific Slender-Toed Geckos, Nactus pelagicus Complex (Reptilia: Squamata), of New Guinea and Adjacent Islands. Smithsonian Contributions to Zoology, (651): x + 92 pages - get paper here
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